ECHEVERIA LISTINGS A-B

By James E. Low

ACUTIFOLIA (Lindley, 1842), Oaxaca state, SW Mexico, 900 ft., species name means: “sharp leafed”, listed in new Series 5 (Gibbiflorae) in the 2003 Lexicon; cytology count still not shown in 2003, thus: n=?: (Uhl found n=28 for his “Ech. aff. acutifolia”). A form found by MacDougall had red leaves (ISI-72), but the usual form has green leaves. See CSJA of 1958, 30(4), 105. Lower stems to 10 cm long, rarely branching, leaves usually green, may be tinged red; the inflorescences cymose, flowers scarlet tinged with yellow. Kimnach calls this an attractive species that is rarely found in cultivation.

=Cotyledon acutifolia [Baker, 1869], obs.

=*Ech. acutifolia [Lindley, 1842], current.

=Ech. holwayi [Rose, 1911], obs.

=Cotyledon devensis [Brown, 1906], obs.

= Ech. devensis [Green. 1931], obs.

adunca [Otto, 18??]. Later found not to be an Echeveria! It is now:

=*Pachyphytum hookeri (moved out of Ech. by Link).

AFFINIS [Walther, 1958], in Durango & Sinaloa states, N. Cent. Mex. [related to Ech. craigiana (thus the Latin species name meaning “related”)]. The wild plants are variable; published in CSSA Journ. 30: 4, 105, 1958. See Journ. CSSA Nov. 1993, p. 272; first listed in old Walther series 1, #6, p.72, (fig.23, p. 73), Uhl moved this from Series Paniculatae to Series Occidentales in 1995 in Haseltonia #3, p. 25, Chromosome numbers are n=30 (diploid) and 60 (tetraploid). Kimnach now lists this in his new Series 9 (Occidentales) in the 2003 Lexicon. Stems slowly branching; medium rosettes with leaf color in wild forms ranging from green to dark, but the most popular form for cultivation was chosen as the darkest leaf form, often nearly black. The wild, green forms are usually larger plants; the inflorescences are flat-topped cymes usually 20 to 30 cm, rarely to 40 cm; flowers red-scarlet.

=*Ech. affinis [Walther, 1958], current.

 

“Ech. cf. affinis” U1097A, (unnamed and unpublished new sp.) Species Nova from NW Mexico in state of Sinaloa; see Haseltonia #3 of 1995 on p. 25 for all details; Series Occidentales, n=30.

AGAVOIDES [Lemaire, 1863], name means “resembling the agave (century) plant”, from states of SLP to Hidalgo; first placed in old Walther series 2, sp. #8, P.81, (fig. 31, p.83), 3 varieties (listed below with + sign). Range: N.E. to E. Cent. Mex.; later in Series Urceolatae, cytology: n=29 & 58. In 2003 Lexicon Kimnach lists this species as being in his new Series 16 (Urbiniae). This is a popular species in nurseries everywhere. The subspecies and forms and cultivars are hard to separate as to origin and classification (as data has been lost over the ages). Beautiful new forms are still being developed or found in nature. Plants are usually unbranched, leaves green sometimes reddened on margins or tip; the infl. is mostly with 2 cincinni, flowers pinkish to orange-red. Plants of easy culture.

=*Ech. agavoides [Lemaire, 1863], current.

=Cotylodon agavoides [Baker, 1869], obs.

=Urbinia agavoides [Rose, 1903], obs.

=Urbinia obscura [Rose, 1903], obs.

=Ech. obscura [Berger, 1930], obs.

=Ech. yuccoides, [Morren, 1874], obs.

+ ‘Prolifera’, [Kimnach 1998], a cultivar, (see CSJA 70(6): 300, ’98). This was a collection of the wild species in a lovely, but extreme form, but not a subspecies (Walther did call it a variety in his book).

=Ech. agavoides var. prolifera [Walther, 1972], obs.

+ ‘Multifida’ [Kimnach, 1998], a cultivar, (see CSJA 70(6); 300, ’98). This is still another collection of a beautiful, but extreme wild form (Walther called it a full variety).

=Ech. agavoides var. multifida [Walther, 1972], obs.

+ ‘Ebony’ [Kimnach & Trager, 1992], a cultivar, (see CSJA 65: 245, 1993). This was another collection of a wild form, thus was technically the full species in one of its extreme forms (found long after Walther had died).

 

+ ‘Corderoyi’ [Kimnach, 1998], a cultivar, (see CSJA 70(6) : 300). This used to be known as various older names listed below. Kimnach recently decided it deserved only cultivar status.

=Cotyledon corderoyi [Baker, 1874], obs.

=Ech. corderoyi [Morren, 1874], obs.

=Urbinia corderoyi [Rose, 1903], obs.

=Ech. agavoides var. corderoyi [Poellnitz,1936], obs.

 

ALATA [Alexander, 1941], Mr. Alexander published field collection number B-16 that was found by MacDougall in SW Mexico, Oaxaca state, 2200 meters alt.; species name means “winged” (for corolla), now in new Series 8 (Nudae) of Kimnach in 2003 Lexicon; cytology: n=17 (read Uhl in AJB, 1982, p. 846), and see CSJA 13: 136, 1941 for illustration. The plant is not attractive, but flowers are (scarlet & yellow), leaves diffuse along stem, no true rosettes, leaves green with red margins; the infl. a medium raceme.

=*Ech. alata [Alexander, 1941], current

 

albicans [Walther, 1958], unknown in nature, (name means “whitewashed”), Walther series 2, #18, p. 110, (fig. 50, p.111), Series 2, n=120-130, now moved to Series Urceolatae, see p. 68 of Haseltonia #4. Read text of Ech. elegans listing below.

=Ech. albicans [Walther, 1958], obs. Published in CSSA Journ. 30:147, 1958. In 2003 Lexicon reduced to synonymy with Ech. elegans

=*Ech. elegans (Rose, 1905), current

alpina [Walther, 1935], from: (see Ech. secunda), name means “mountain plant”, published in CSSA Journal 7:70, 1935; first in old Walther series 2, sp. #34, p. 146, reduced to synonymy with Ech. secunda by Uhl on p. 35 of Haseltonia #3 in 1995, n=30, 32.

=Ech. alpina [Walther, 1935], obs.

=*Ech. secunda [Uhl, 1995], current.

AMOENA (De Smet, 1875), means “charming”, E. Cent. Mex. in states of Vera Cruz & Puebla, found near Vera Cruz city on limestone. This is a small, pretty plant, freely offsetting & the tiny leaf-like bracts between flowers drop easily and root to make new plants; flowers salmon-orange color. Walther series 1, #3, p.67; this species was in earlier Series 1, but now is moved into Kimnach’s new Series 10 in 2003 Lexicon; cytology: n=33, 66. Rosette leaves are glaucous-green, often red tipped, the infl. a lax cyme, flowers salmon-orange. The 3 older names combined below exist as distinct forms today.

= Ech. pusilla [Berger, 1904], obs.

=Ech. purpusii [Britton, 1905], obs.

=Ech. microcalyx [Britton & Rose, 1911], obs.

=*E. amoena [De Smet, 1875], current.

 

 

 

 

AMPHORALIS [Walther, 1958], SW Mexico in state of Oaxaca growing at 7000 ft.; name means the flowers “having shape of a flask”; published in CSSA Journal, 30:149, 1958; now in new Series 4 (Echeveria) in 2003 Lexicon; cytology: n=38, MacDougal field number B-82. Leaves, etc. are all puberulous-hairy, leaves green with red margins. Infl. to 20 cm. Petals scarlet-red with yellow margins.

=*Ech. amphoralis [Walther, 1958], current.

ANDICOLA [Pino, 2005], of central Peru in Dept. Junin, Pasco, & Huanuco at around 3170 meters growing on slopes in clay between rocks. (Series ? not stated, but probably in Racemosae, series 12, like its two relatives E. chiclensis & E. peruviana.) Chromosome count not yet taken: n=? Plant: single medium rosette at ground level, leaves light green to pink blushed. Infl. a medium spike with urn shaped flowers of bright pink. Name andicola means “living in the Andes mts.”. Species published in Haseltonia #11 (2005) pages 98-102. This species was at first confused with species E. chiclensis or E. peruviana which it closely resembles, but this grows at lower altitudes. It is fairly common over a large range, and is variable in leaf form over this range. Its closest related species seems to be Ech. excelsa of the same area, but excelsa is much larger and flowers are red.

=*Ech. andicola [Pino, 2005], current.

ANGUSTIFOLIA [Walther, 1972], NE Mexico (SLP state ?), name means “narrow leafed”; species published by Walther in his 1972 book ‘Echeveria’ on p. 211; no live plants of type-species exist today: chromosome count for original plant is thus unknown, so cytology: n=? See Haseltonia #4 of 1996 on p. 68 and 77 for a fuller discussion, in 1996 tests were carried 2 plants field collected by Uhl called: “cf. angustifolia”. Prof. Uhl called these “Ech. affiliated with Ech. angustifolia” but it is possible they are the same. He then tested both plants and found the chromosome count to be n=ca. 220; so he concluded they are probably in his Series Urceolatae. Later Kimnach accepted (same?) 2 plants (of Tam. & SLP states) as valid and placed the species in his new series 16 (Racemosae), in the 2003 Lexicon, also accepting the high chr. counts & this description: plants not branching, the inflorescence a 20 cm tall, single cincinnus with 10 to 15 fl.; the flower was reddish. Kimnach noted this plant resembles Ech. bifida (with n=12), but these 2 new plants had the much higher chromosome (polyploid) count.

=*E. angustifolia [Walther, 1972], current.

ATROPURPUREA (Morren, 1874), name means (having leaves of) “ugly purple”, possibly from Vera Cruz, Mex. Published in Belg. Hort. 156, 1874. Original plants lost: some new plants seem valid, but habitat for them unknown. This species now placed in new Series 12 in 2003 Lexicon, but no cytology count shown, thus n=?. New plants form easily from dropped bracts: mother plants rarely branch and have 15 cm main stems below rosettes. Leaves deeply channeled, reddish-brown; flowers red and in medium tall racemes.

=Cotyledon atropurpurea (Baker, 1870), obs.

=*E. atropurpurea [Baker, 1972], current.

 

AUSTRALIS (Rose, 1903), found widely over Central America, usually growing on trees (epiphytic), name means “southern” type; now placed in new Series 8 (Nudae) in 2003 Lexicon; cytology: n=28. Plants are “sub-shrubs”, main stems to 30 cm tall. Leaves hairless, light green, glaucous, often tinged purple. Flowers in dense racemes, corollas angular and red. Plants are highly cold sensitive.

=*E. australis [Rose, 1903], published in Bulletin of NY Bot. Gard. III:6, 1903.

 

backebergii (Poellnitz, 1935), from E. Cent. Peru; published in Fedde Rep. 1935, 38:185, name honors German botanist Curt Backeberg; see Hasetltonia #5 on p. 51.

=Ech. backebergii [Poellnitz, 1935], obs.

=*E. chiclensis var. backebergii [Pino, 2002], current. See details in this new listing.

BAKERI (Kimnach, 1991), from Bolivia, South America, grows at 11,000 ft.; name honors Wm. Baker, species published in CSSA Journal, Sept. 1991, p. 254; now in new Series 12 (Racemosae) in 2003 Lexicon, chromosome count: n=160; main stems 10 cm, rosettes usually single, leaves glaucous to reddish; flowering stems tall racemes: 50 to 90 cm, flowers pinkish-orange. (#) Color photo on p. 198 of CSSA Journal of Sept. 2007.

=*E. bakeri [Kimnach, 1991], current.

BALLSII (Walther, 1958), from Columbia, South America, species published in CSSA Journal, 30(2):44, 1958. Now in new Series 12 (Racemosae) in 2003 Lexicon, cytology: n= approx. 40. Name honors early botanist Ball; type sp. was his field number: Ball-7587. Main stems short, but branching; leaves green, inflorescences racemes 25 to 30 cm tall with about 10 nodding flowers; the flowers orange-red to scarlet.

=*Ech. ballsii [Walther, 1958], current.

BELLA (Alexander, 1941), S. Mexico in state of Chiapas, species discovered by MacDougall & published in CSSA Journal 13:133,1941; name means “beautiful”. Now in new Series 12 (Racemosae) as listed in 2003 Lexicon, cytology: n=15. Species variable as found in nature so there are some recognized forms, but none are still called varities. Main stems short but heavily branching with small rosettes of plain green leaves; inflorescences are racemes to 25 cm tall; flowers coral with yellow at apex. This species considered to be free flowering, but difficult in cultivation. The fma. major is similar but lager.

=*Ech. bella [Alexander, 1941], current.

+*Ech. bella fma. bella [Walther 1972], current.

+var. major (Walther, 1972), obs. This now becomes:

=*Ech. bella fma. major [Kimnach, 1997], current. See details in Haseltonia #5 on p. 51.

BICOLOR (Knuth, 1823; Walther, 1935], of S. America (Venezuela to Columbia), name means “having 2 colors” (in flowers); leaves of type-species plain green, others may be green with some glaucous coating. As found in nature this species is extremely variable over its rather large range. Check the following references:

CSSA Journ. 1935, 7:39, (republished as an Echeveria by Walther).

" " 1991, Mar. p.25, (ISI 91-42). Plant offering.

" " 1992, Jan. p.3, by Charles Uhl with photos.

" " 1993, Mar. p.81 by Uhl (forms #1, 2, & 3 described)

First placed in old Series Racemosae, now in new Series 8 (Nudae) in 2003 Lexicon; cytology: n= 21 (diploid), and rarely n=42 (tetraploid). Brief description: main stems go to 50 cm tall or more, but are little branched. The inflorescences are racemes with reddish-yellow flowers. Species discovered long ago so there are several old names (some in wrong genus):

=Sedum bicolor [Humboldt, Bonpland & Knuth, 1823], obs. H&B discovered it, Knuth published it.

=Cotyledon bracteolata [Link, K. & O., 1869], obs.

=Ech. bracteolata [Link, Klotzsch, & Otto, 1842], obs. (not proven, but Uhl thinks probably same).

=Cotyledon subspicata [Baker, 1869], obs.

=Ech. subspicata [Berger, 1930], obsolete for now; but more study may return this more western type to full species status (it is still little known).

=Ech. bicolor var. subspicata [Walther, 1935], obs.

=Ech. venezuelensis [Rose, 1930], obs. see Uhl in 1992 text (above).

=*Ech. bicolor [Walther, 1935], current. This is the type-species: flowers of yellowish to reddish colors on taller inflorescences.

+*Ech. bicolor var. turumiquirensis [Steyermark, 1952], current. (from E. Venez. in Sucre & Monagas, see CSSA Jan.’92). Differs in having shorter infl. stems & flowers deep red.)

 

 

BIFIDA (Schlechtendal, 1839), E. Cent. Mex. state of Hidalgo & maybe even in Queretaro; Latin name means “split into”, published in Linnea XIII: 411, 1839, (see CSSA Journ. Mar. 1976, for ISI-963 offering), now in new Series 1 (Angulatae) in the 2003 Lexicon, cytology: n=12. Main stem to 6 cm by 1 to 1.5 cm and usually unbranched; leaves green to purplish; infl. 3 branched; flowers reddish & yellow. Note: this may be same species as the little known Ech. teretifolia of De Candolle, now ignored (Kimnach explained this at length in the new Lexicon under name bifida).

=Cotyledon bifida [Hemsley, 1880], obs.

=Ech. bifurcata [Rose, 1909], obs.

=Ech. erubescens [Walther, 1972]. obs.

=Ech. tenuifolia [Walther, 1972], obs.

=*Ech. bifida [Schlechtendal, 1839], current.

 

 

bifurcata, (Rose, 1909], E. Cent. Mex., means (inflorescence is) “double forked”, refer to Haseltonia #6., p. 65 & 73. Published in Contr. US Nat. Herb. 12:439, 1909. Note: this name has now been combined into bifida (see above).

=Ech. bifurcata [Rose, 1909], obs.

=*Ech. bifida [Schlechtendal, 1839], current.

 

BRACHETII (Reyes & González-Zorzano, 2009) published in Cactáceas y Suculentas Mexicanas, 54(3) : 78, ills., 2009. Found in the Sierra de Juárez, Oaxaca, Mexico, growing in pine-oak forests and prefering sites with high humidity, enough organic matter and some shade.
This is a glabrous plant, caespitose, stems up to 30 cm tall, rosettes loose, 5 – 9 cm in diameter; leaves glaucous, obovate-spatulate, mucronate, colour light green; inflorescence 20 – 30 cm long, upright or decumbent, with ca 14 flowers; pedicels up to 10 mm long, sepals ascending, corolla up to 10 mm long, orange to pink, petals inside yellow.
Placed in Series Nudae because of the conspicuous stem though its inflorescence is different.
=*Ech. brachetii [Reyes & Gonzáles-Zorzano, 2009] current.

 

bracteolata (Link, Klotzsch, & Otto, 1844), Venezuela, mentioned in CSSA Journ. Jan. 1992, p. 3, & in issue of Mar.1993, p. 74; see also CSSA Journ. of Jan. 1992, p. 3. (bracteolata name refers to having bract-leaves between flowers); this is now combined into Ech. bicolor.

=Ech. bracteolata, [L, K & O, 1844], obs.

=*Ech. bicolor, [H, B & K, 1823; Walther, 1935], current.

brittonii (N.& M., 1913), this species now known to be a Sedum. Northern Calif. high altitudes.

=Ech. brittonii [N&M, 1913], obs.

=Sedum hallii, [Praeger, 1921], obs.

=*Sedum obtusatum, [Gray, 1868], current.

buchtienii (Poellnitz, 1934), Bolivia: now combined into Ech. whitei: see that listing for all details.

=Ech. buchtienii, [Poellnitz, 1934], obs.

=*Ech. whitei (Rose, 1925), current status.

byrnesii (Rose, 1905); this name was informally reduced to synonomy with Ech. secunda by Uhl on p. 35 of Haseltonia #3 of 1995, his action formalized by Kimnach later, see below.

=Ech. byrnesii [Rose, 1905], obs.

=*Ech. secunda (Kimnach, 1997), current. (Comb. nov. in Haseltonia #5, p. 51).