ECHEVERIA LISTINGS C

By James E. Low

CALDERONIAE (Perez, 1997), of central Mexico state of Guanajuato; species name honors Mrs. Graciela Calderon de Rzedowski (a Mex. botanist); first placed in old series Secundae, but now in new Series 13 of same name in 2003 Lexicon; chromosome count still not given in new Lexicon, thus: n=? Published in Acta Botonica Mexicana of 1997, including a line drawing showing a plant with many long, narrow, pointed leaves in rosette, and a medium long flowering spike with few flowers (about 5, but up to 20 seen elsewhere), fl. orange to reddish, yellow inside, elongated. (Such linear leaves unique to Secundae Series). Type plant found 4 or 5 km south of Santa Barbara in the village of Ocampo at an altitude of 2200 meters growing under live oaks in 1995. Stems very short, but branching; leaves flat on top, and convex below are about 5 cm long and curve a bit upward. Rosettes single or clumping, 20 to 30 green leaves, total dia. of 12 cm .
=*Ech. calderoniae [Perez. 1997], current.

CALYCOSA (Moran, 1966), of Cent. Mexico state of Michoacan, name means having a “prominent calyx”, species published CSSA Journ. 38:14, 1966; plants offered in CSSA Journ. Mar. 1976, p. 90 as ISI-964, first placed in old Series Valvatae, now in new Series 17 of same name; cytology: n=31: see Haseltonia #5, p.22 & 31, rosettes single, infl. a cincinnus to 21 cm; flowers yellow & orange, leaves light green. This plant delicate & difficult in cultivation. =*Ech. calycosa [Moran, 1966], current.

CANALICULATA (Hooker, 1857), S. Mex., from Chiapas to Oaxaca state, name means “having channels” (in leaves); species published in Curt. Bot. Mag. of 1847; plants offered in CSSA Journ. Mar. '80, p.101 as ISI-1196, now in new Series 12 (Racemosae) in 2003 Lexicon, cytology: n=22 or 44 (new Lexicon gives only 44). Main stems to 15 cm (branching ?); leaves deeply channeled, glaucous tinged purple (or light green farther to the south in range); infl. are long racemes 35 to 50 cm; flowers brick red. Original plants lost, new plants from Chiapas unverified. Plants from cultivation with this name have green leaves, small corollas & habitat uncertain.
= (?) canaliculata [Hooker, 1857], obs.
=Cotyledon canaliculata [Baker, 1869], obs.
=Ech. rubescens [Lemaire, 1857], obs.
=*Ech. canaliculata [Hooker, 1857], current.

CANTE (Glass & Mendoza, 1997), of central Mex., a long known but improperly named species, related to Ech. subrigida; name ‘cante’ is American Indian for “life-water”; published in CSSA Journ. Sept. 1997; now in new Series 5 (Gibbiflorae) in 2003 Lexicon; cytology: n=27(?), main stem to 10 cm tall, not branching, rosettes quite wide to 40 cm. dia, leaves heavily white frosted (pruinose) tinged lavender with reddish margin, inflorescences cymose, from 45 to 60 cm tall and pruinose, flowers orange-pink. This species was long confused with Ech. subrigida which has several minor details of difference.
=*Ech. cante [Glass & Mendoza, 1997], current. = “false Ech. subrigida of Zacatecas” (name used by some growers since 1930s).

CARMINEA (Alexander, 1941), SW Mex. in Oaxaca state, name means “carmine”, species published in CSSA Journ. 13:138, 1941, first in old Series 11 (Racemosae), now in new Series 4 (Echeveria) in 2003 Lexicon; Cytology: n=21; species discovered by MacDougall. Stems can become tall to 70 cm. with loose rosettes, becoming a small shrub; leaves and stems hairy (finely pubescent), green, margins red; flowers large and showy, orange with yellow; or rarely just yellow.
=*Ech. carminea, [Alexander, 1941], current.

Echeveria carminea, Kimnach 88047, Oaxaca, Quiechapa, Maderera, Lau 087, Hbg 41270 (Photo Helmut Regnat)

CARNICOLOR (Morren, 1874), E. Cent. Mex., Vera Cruz, name means “meat colored”, species published in La Belg. Hort. 24: 158, 1874; now in new Series 12 (Racemosae) in 2003 Lexicon, cytology: n=18. Plants branching, medium rosettes; leaves densely papillose, glistening nicely, grayish to purplish-green leaves; short flowering stems to 15 cm, a 2-sided racemose, bract leaves fall and root easily; flowers salmon to flesh colored. Plant resembles Ech. waltheri, but differs mostly in its chromosome count.
=Cotyledon carnicolor [Baker, 1870] obs.
=*Ech. carnicolor [Morren, 1874], current.

CHAPALENSIS (Moran & Uhl, 1989), W. Cent. Mex., Jalisco state, near Lake Chalapa (thus the name) at 5500 ft., species published in Mex. C&S Journ. XXXIV, 2:27, 1989; now in new Series 8 (Nudae), cytology: n=45. Main stems to 50 cm tall, branching, small rosettes, leaves finely muriculate-papillose, green with red margins; inflorescences spicate 20 to 50 cm tall, flowers on the long spike are greenish to rose above.
=*Ech. chapalensis [Moran & Uhl, 1989], current.

CHAZAROI (Kimnach, 1995) of SW Mex. state of Oaxaca, 6400 ft. elev., honors Prof. Miguel Chazaro of Univ. of Guadalajara, species published in CSSA Journ. Mar. 1995, p. 83, was first in old Series Racemosae; now in new Series 12 of same name in 2003 Lexicon, chromosome count still not given in new 2003 Lexicon, thus: n=? Stems rarely branching, medium rosettes, leaves are bluish-green with paler edge band. Inflorescences are racemose to about 45 cm tall, flowers salmon-pink. This species may not now be in cultivation: could be extinct soon after discovery. Color photo on p. 198 of CSSA Journ.of Sept. 2007.
=*Ech. chazaroi [Kimnach, 1995], current.

chiapensis [Rose, & Poellnitz, 1936], E. Cent. to SW Mex. in States of Vera Cruz, Puebla, Chiapas, & Oaxaca, 2300 M. elev., name means “of Chiapas”, this species was combined into Ech. rosea of Lindley in the 2003 Lexicon.
=Ech. chiapensis [Rose & Poellnitz, 1936], obs.
=*Ech. rosea, [Lindley, 1842], current.

CHICLENSIS (Berger, 1930), of Peru, S.A., near Chicla (thus the name), first placed in old series Racemosae; now in new Series 12 of same name in 2003 Lexicon; cytology: n= approximately 120. Published in Engl. Nat. Pflzfam. 1930. II:18a:473. This species has unusual roots (tuberous), and it is said to be a difficult plant in cultivation. Wild pants quite variable: some may be very hairy, others with no hair (the one with longest hair is now a valid form: see below). Roots thick fusiform, stem short, rosettes (?), leaves green usually hairless, the inflorescences are racemes to 30 cm tall; flowers yellowish & pinkish according to local form.
=Cotyledon chiclensis [Ball, 1887], obs.
=Ech. neglecta [Poellnitz, 1935] ), obs.
=*Ech. chiclensis, [Berger, 1930], current.
=*Ech. chiclensis var. chiclensis [Pino, 2002], current: the type species.
+Ech. backebergii [Poellnitz, 1935], obs. This becomes the form shown below:
=Ech. chiclensis fa. backebergii [Kimnach, 1998], Comb. nov. Haseltonia #5, p. 51, obs.
=*Ech. chiclensis var. backebergii [Pino], Haseltonia #9. This is the very hairy form in which the hairs glisten in the sun.

CHIHUAHUAENSIS [Poellnitz, 1935], N. Cent. Mex., states of Chihuahua, Sonora & Durango; species published in Fedde. Rep. 38:29, 1935; see CSSA Journ Sep. '93, p. 245, and CSSA Mar. '80, name means “of Chichuahua” (state), plants rarely branching, leaves light blue-green and coated glaucous white (rarely dark green in nature) tips red; free flowering; inflorescence: a 1 to 5 branched cincinnus about 20 cm tall, flowers reddish. First placed in Walther’s old series 2 as sp. #10 on p. 88, Uhl reviewed it in Haseltonia #4 of 1966 and keeps it in Series Urceolatae (on p. 66 & 77 of that issue), but in 2003 Lexicon Kimnach places it in his new Series 16 (Urbiniae); with cytology of n=26, & 50.

=*Ech. chihuahuaensis [Poellnitz, 1935], current.

CHILONENSIS (Walther, 1935), Bolivia, South America near Sucre at 3000 M. elev., Dept. of Chuquisaca; published in CSSA Journ, 1935, 7:40; now listed in new Series 12 (Racemosae) in 2003 Lexicon; chromosome count difficult to read, but approximately n=165. Medium rosettes, leaves reddish green to plain green, inflorescences a long 60 cm raceme, flowers nearly white or yellowish.
=Sedum chilonense [Knutze, 1898], obs.
=Ech. vanvlietii [van Keppel, 1969], obs.
=*Ech. chilonensis [Walther, 1935], current.

ciliata (Moran, 1961) of S. Mexico in NW Oaxaca state. Published in CSSA Journl 13:131, 1961; Moran downgraded this from full species in CSSA Journ. p. 27-36, Jan. 1993; see photo, fig.14, p. 99 CSSA Journ. of Mar. 1995, see also AJB, 1976, p. 810, & AJB 1992, p. 843-854.
=Ech. ciliata [Moran, 1961], obs.
=*Ech. setosa var. cilata (Moran, 1963

COCCINEA (De Candolle, 1828), E. Cent. to SW Mex. states of Hidalgo, Fed. Dist., Puebla, to Oaxaca, name means “scarlet”, see CSSA Journ. Sep. '76, p. 225; was first in old series 13, but now in new Series 4 (Echeveria) in 2003 Lexicon, cytology: n=23, 25. Read history of this species page 303 of CSSA Journ. of Nov. 1999 with early painting. This was the first Echeveria published after this genus name was established. Leaves are covered with velvety white pubescent hairs, surface green to reddish under the white velvet. Most parts of plant are pubescent except inside flowers. Lower stems to 60 cm tall, sparsely branching, rosettes loose, leaves green or white hirstute; the inflorecences are spikes up to 70 cm tall, scarlet flowers. Some plants in the wild have wide leaves and were called Ech. pubescens, but this is a spectrum and old pubescens was just a part of it, and thus cannot be retained.
=Sedum spicatum (Sesse & Mocino, 17??), obs. (never formally published).
=Cotyledon coccinea (Cavanilles, 1793), obs.
=Cotyledon pubescens (Schlechtendal, 1839) for wide leaf type, but incorrect genus. Obs. It became:
=Ech. pubescens (Baker, 1869), obs.
=*Ech. coccinea (DC, 1828), current.

C'est la plante qui autrefois était connue sous le nom de E. pubescens. Elle a été inclue dans E. coccinea parce que la forme des feuilles de la dernière est fort variable - des feuilles étroites et des feuilles très larges et beaucoup de formes de transition, même des formes aux poils blancs.

This is the plant formerly called E. pubescens. It has been included in E. coccinea because the leaves of the latter are very variable - leaves narrow or wide and transitional forms as well as leaves with white hairs.

colomuiana, ? (mentioned in Haseltonia #3, p. 28, 1995 by Uhl as a collected plant he tested as having n=44, and as being in Series Racemosae). No further info.

COLORATA (Walther, 1972), W. Cent. Mex. states of Jalisco & Guadelajara at 9,500 ft. alt., published in Walther’s book ‘Echeveria” in 1972, first placed in old Walther series 2, #2, p. 91, but in 2003 Lexicon now listed in new Series 16 (Urbiniae). It was ISI offering, p. 101 (ISI 1198), Mar. 1989, p. 59 for ISI 89-49. Species name means “colored” (for leaf margins): the red of leaf edges may be due to a virus! (closest relative Ech. chihuahuaensis), the chromosome count: n=27. In 1980 Kimnach gave a form (brandtii) full variety status, but downgraded this later to mere forma when it was found to be just a part of a spectrum of forms. This also happened to the other well known form that for a time was called ‘Ech. lindsayana’. A full “spectrum” of types are indeed found in nature, and some are very nice and worth collecting. Most are rather large plants, rarely branching, leaves are blueish-glaucous suffused red, inflorescences 30 cm tall, 2 to 3 branching, each cincinni 5 cm with 4 to 6 flowers, flowers reddish or orange at base shading to yellowish. Color photo of this “Mexican Giant” on p. 200 of CSSA Journ. of Sept. 2007.
=*Ech. colorata [Walther, 1972], current.
=*Ech. colorata forma colorata [Kimnach, 1980], current for type species.
+*Ech. colorata forma brandtii [Kimnach, 1980], current for this one variety.
=Ech. colorata var. brandtii [Kimnach, 1980], obs
=Ech. lindsayana [Walther, 1972], obs.

COMPRESSICAULIS (Eggli & Taylor, 2002) Venezuela, 25 mi. S. of Merida at 6,500 ft. elev. Taylor collection #675 (1988), ISI 2002-20, HBG 75704, Kew 485-88-03745. Published in Curtis Botanical Mag. 2002, 19(4):222-226. Placed into the new Series 12 (Racmosae) in the 2003 Lexicon. No chromosome count given in Lexicon account, but elsewhere is given as n=42 (tetraploid). Plants upright growing to about 5 in., base branching. The stems show “compression” scars prominently, thus the Latin name meaning “compressed stem”. Rosettes minimal: leaves loose and scattered & of a distinctive purplish-brown color over green. The inflorescence a 2-sided, 20 cm raceme with stem also showing compression marks. Flowers orange, campanulate. Its odd stems resemble E. nodulosa of Mex. its closest relative, but it seems mysterious that their habitats could be so far apart. This plant was long known and in cultivation widely, but it remained unnamed until now, as it seemed to be part of a “spectrum” of types, thus hard to decide where to draw a line for distinct species, this plant mentioned by Uhl on p. 200 of the July, 1992 issue of CSSA Journ. This species probably related to Ech. pendulosa of same area, and with same chr. count. (but its leaves: blue-grn.) Compressicaulis is said to be of easy culture, and easy to propagate from leaves.
=*Ech. compressicaulis [Eggli & Taylor,2002], current.

corallina [Alexander, 1941], found to be same as “Ech. sessiliflora”, thus now combined into it.
=*Ech. sessiliflora [Rose, 1905], current.

cornuta (Walther, 1972), E. Cent. Mex. Hidalgo state near Jalisca; name means “horned”, reduced to synonomy with Ech. secunda by Uhl on p. 35 of Haseltonia #3 of 199; see that species listing for all details.
=Ech. cornuta [Walther, 1972], obs.
=*Ech. secunda [Booth, 1838], current.

courantia echeverioides, [Lemaire, 1851] is now reclassified as Echeveria rosea: see that listing for details. Published in Bot. Reg. 1842, 28:22. Courantia is a discontinued genus name.
=C. echeverioides [Lemaire, 1851], obs.
=C. rosea [Lemaire, 1853], obs.
=*Ech. rosea [Lindley, 1842], current.

CRAIGIANA (Walther, 1952), N. Cent. Mex. in states of Chihuahua & Sonora, published in CSSA Journ. 24(1):28, 1952. First placed in old Walther series 1, sp. #7, p. 76 (fig. 8, p. 77), read CSSA Journ. p. 57 of Mar., 1980 and issue of Jul. 1990, p. 172, later moved by Moran in 1977 to his Series 9 (Occidentales), and today appears in 2003 Lexicon in new series 9 of same name; the cytology is n=30 (diploid). The species name honors Rob. Craig (see p. 57, CSSA, Mar. 1980). 2 inch stem, rarely branching; leaves brownish-green, slightly glaucous; cymose inflorescences to 50 cm tall, flowers red, n=30.
=*Ech. craigiana (Walther, 1952), current.

crassicaulis (Walther, 1935), E. Cent. Mex. states of Puebla, Fed. Dist., & Vera Cruz, species published in CSSA Journ., 7:36, 1935; name means “hard stemmed”, (Prof. Uhl thinks this sp. and Ech. platyphylla, plus Ech. mucronata may all be combined into a single species in time); Note; in the 2003 Lexicon E. crassicaulis was combined only with E. mucronata under that name. Is now in new Series 7, (Mucronatae), cytology is n=16, 32.
=Ech. crassicaulis [Walther, 1935], obs.
=*Ech. mucronata [Schl. 1839

CRENULATA (Rose, 1911), E. Cent. Mex. in Morelos state near Cuernavaca, collected by Rose & Painter #797; species published in Contr. US Nat. Herb., 1911, XIII:295; name means “notched” referring to leaf edges, but in reality these are finely undulate, not much crenulate; first in old Walther series 2, #54, p.196, now in new Series 5 (Gibbiflorae) in 2003 Lexicon; Chromosome count not given in the 2003 Lexicon, thus: n=? Base stems to 5 inches, rosettes (?) and branching (?); leaves plain green, not glaucous and with margins dark red-brown; inflorescence cymose to 50 cm tall with 9 or more cincinni with 4 to 12 flowers ea. Flowers pink to yellowish.
=*Ech. crenulata [Rose, 1911], current.

CUENCAENSIS (Poellnitz, 1935), of Equador, S. A. at Cuenca (species name means "of Cuenca"), species published in Fedde Rep. 1935, 38:187; now in new Series 9 (Racemosae) in 2003 Lexicon; cytology: n=approx. 100; the stems unknown, leaves grey-green with reddish margins; inf. 20 cm racemes with about 30 fl. red to yellowish. Not today in cultivation: this is still a little-known species and growth habit is unknown in 2003, but Walther in 1972 wrote that he thought it related to E. excelsa of Peru. This species needs more research.
= Ech. cuencaensis [Poellnitz, 1935].
=*Ech. quitensis [Harling & Andersson, 2004] current.

CUICATECANA [RSJ, etc., 2004], of southern Mexico, state of Oaxaca at Cuicatlan (thus the Latin species name), growing in tropical deciduous forest by a river, plants hanging pendant in large clusters on vertical slope. This is a new species similar in appearance to E. lauii, but with curious differences. This species is tentatively placed in Series Pruinosae, but the floral structure and chromosome counts are not typical of that series. Chromosome count is n=30. Flowers bright pink, but almost hidden under large, succulent, white sepals. The flower cluster spikes arch downward. These parts closely resemble those of a Pachyphytum to which it may be related. [Note that Kimnach (2004) seems to think it should be classified as being in genus Pachyphytum, so the matter may not yet be settled]. The 3 publication authors (shown only as “RSJ, etc.” above) list their names in the Spanish way, thus: “Reyes Santiago, Jeronimo”, and “Perez Cristano, Joel”, and “Brachet Ize, Christian”. They work for the botanical garden in Mexico City. Publication in the Mexican cactus and succ. Journal XLIX, 49(3) on pages 80-84, year 2004.
=*Ech. cuicatecana [Santiago, Cristano, & Ize; 2004], current (?!).

CUSPIDATA (Rose, 1905), NE to N. Cent. Mex. This is a saxicolous (rock dwelling) species, and is now known to exist in 3 varities in ranges near Saltillo, Coah.; its Latin name means it has “pointed” leaf tips, species published in Bulletin NY Bot. Gard. 1903, III:9. First placed in old Walther series 2, as species #32. p. 144, but now in new Series 16 (Urbiniae): see 2003 Lexicon. See Haseltonia #4 of 1996, page 68. where Moran, Kimnach & Uhl moved this sp. from Walther’s Series 2 (Secundae) into Series 3, (Urceolatae), but in new Lexicon Kimnach has it in his new Series 16; cytology given as n=31. Note that a full history of this species is presented by Kimnach in his article where he published the 2 new subspecies. This was in the CSSA Journ. of Jan. 2005 starting on p. 28 to 33 (+p. 47). For type species: main stems thick and to 5 inches long, usually un-branched, rosettes 6 to 22 cm in dia., leaves gray-glaucous tinged reddish, the inflorescences 40 cm with 6 to 16 fl. The flowers scarlet, tinged orange. Walther thought E. parrasensis to be a separate species, but more recent study showed it to be merely a smaller form of E. cuspidata. Color photo on p. 199 of CSSA Journ. of Sept. 2007.
=Ech. parrasensis (Walther, 1972) obs.
=*Ech. cuspidata (Rose, 1905), current for type sp.
+ 2 new subspecies in 2005 in CSSA, p. 31 & 33:
=Ech. cuspidata var. zaragozae (Kimnach, 2005) range: NL state on Gypsum hills 10 mi. south of Aramberri. Current.
=Ech. cuspidata var. gemmula (Kimnach, 2005), range: NL state on gypsum hills in canyons between towns of Zaragoza & Aramberri at 1200 m altitude. Name means “little gem”. Current