ECHEVERIA LISTINGS D-E-F

By James E. Low

DACTYLIFERA (Walther, 1972), NW Mex., states of Jalisco, Sinaloa, Durango, & Aguascalientes; name means “finger bearing” (see below), first in Walther old series 2, #48, p. 179, but in 2003 Lexicon it is in new Series 5 (Gibbiflorae). Species first seen in 1897, but not pub. until 1972 by Walther on p. 179 of his book, cytology: n=54. Has large rosette on short stems, unbranching, leaves reddish-brown, slightly glaucous; inflorescence cymose to one meter tall! Flowers pink below to orange above, and they have a strange fingerlike “process” near the base, thus this odd Latin species name. An attractive and imposing plant when well grown.
=*Ech. dactylifera [Walther, 1972], current.

DECUMBENS (Kimnach, 1995) from S. America, Peru near Cuzco at 10500 ft.; pub. in CSSA Journ. Jan., 1995, p.3; a dwarf sp. with decumbent flowering stem (thus species name meaning “decumbent”), growth habit similar to that of Ech. westii but most closely related to Ech. peruviana, now placed in the new Series 12 (Racemosae) in 2003 Lexicon, cytology: n=175 (+ or -5). Globose rosettes atop 2 inch, branching stems. Leaves light green to olive-tan, inflorescence a horizontal raceme with ascending portion 26 to 46 cm long, flowers yellowish to red below.
=*Ech. decumbens [Kimnach, 1995], current.

DERENBERGII (Purpus, 1921), SW Mex. state of Oaxaca in Mtns. Mexteca range, pub. in Mosatsschr. Kakteenk, 1921, 31:8, see CSSA Journ. Jul. '94, p. 172, name honors a German med. Dr. Derenberg who loved succulents; now in new Series 3 (Ciliatae) in 2003 Lexicon, cytology: n=27, see Haseltonia #5, p. 22 & 23. The attractive rosettes on short, heavily branching stems, leaves light green, glaucous, tips and margins red; 4 or more cincinni to 10 cm tall; erect, pure yellow flowers. This is a popular succulent in nurseries everywhere, easy culture. It is also useful for making hybrids.
=*Ech. derenbergii [Purpus, 1921], current.

desmetiana (Morren, 1874), name honors Belgian nurseryman & horticulturist Louis De Smet (1810-18??) who published Ech. amoena in 1875.
=Ech. desmetiana [Morren, 1874], obs.
=Ech. subsessilis [Rose, 1905], obs.
=*Ech. peacockii [Croucher, 1874], current.

DIFFRACTENS (Kimnach & Lau, 1981), E. Cent. Mexico near Vera Cruz; name means: easily “shattered” (detached) bract leaves, a 1978 discovery of Alfred Lau (Lau-70), pub. CSSA Journ. Jan. 1981, p.4, BW photo CSSA of May 1991 p.116, ISI offering CSSA Mar. '81 (ISI-1258), related to Ech. carnicolor, now in new Series 12 (Racemosae), cytology: n=18; medium rosettes on 2 inch stems, moderately branching; leaves described by Kimnach as “light pinkish-grey-violet”, inflorescences 1 to 3 branched racemes, flowers orange to yellow at apex. This sp. allied to E. carnicolor. Color photo on p. 199 of CSSA Journ. of Sept. 2007.
=*Ech. diffractens [Kimnach & Lau, 1981], current.

elatior (Walther, 1935), E. Cent. Mexico in Hidalgo state, Walther series 2, #26, p.131, (name may mean tall), pub. CSSA Journ. '35, 7:72; reduced to synonomy with Ech. secunda by Uhl on p. 35 of Haseltonia #3 of 1995; see that listing for all details. This is just a form in the spectrum of types of the common species E. secunda, so it could not be kept as a full species.
=Ech. elatior [Walther, 1935], obs.
=*Ech. secunda (Uhl, 1995), current.

ELEGANS (Rose, 1905), E. Cent. Mex. state of Hidalgo (& most adjoining states) in mtns., name means “elegant”, Walther series 2, #15, p.97, (fig. 39, p.98), see p. 80 in Haseltonia #4, 1996 for Uhl’s discussion, old Series 2, later in revised Series Urceolatae, but now in new Series 16, (Urbiniae) in 2003 Lexicon; cytology: n=31, 32, 34, 60, 62-63, 96, 120-130. Plants have little main stem, rosettes branch freely, leaves in lovely and dense rosettes of greenish-glaucous, to white color. Inflorescences are a cincinnus 10 to 15 cm tall, flowers are rose to pink shading to yellow-green at apex. This species has long been one of the most popular with collectors and in nurseries and gardens. It is a bit more frost sensitive than most. It is a bit variable in nature, and collected extreme forms were published as full species or as varieties (see below), but these were too weak botanically to stand, so were combined by Kimnach in 2003 Lexicon. Color photo on p. 200 of CSSA Journ. of Sept. 2007.
=*Ech. elegans [Rose, 1905] current
=Ech. perelegans [Berger, 1930] obs. Just Berger’s proposed name.
=Ech. albicans (Walther, 1958) obs. Just a whiter form.
=Ech potosina (Walther, 1935) obs. Just a slightly different form.
=Ech. hyalina (Walther, 1958) obs. “ “ “
+var. elegans (Rose, 1905), obs. Walther’s #15a, n=62-63.
+var. hernandonis (Walther, 1972), obs. Walther’s #15b, n=96.
+var. tuxpanensis [Walther, 1972], obs. Walther’s #15c.
+var. simulans (Poellnitz, 1936), obs. Walther’s #15d.
+var. kesselringiana (Poellnitz, 1936) obs.

erubescens (Walther, 1972), possibly from Cent. Mex. state of Queretaro near Bernal, means “becoming red” (when leaves exposed to sun), pub. in Walther’s book ‘Echeveria’ in 1972, p. 263; also see p. 64 & 73 in Haseltonia #6.
=Ech. erubescens [Walther, 1972], obs.
=*Ech. bifida [Schl. 1839], current.

EURYCHLAMIS (Berger, 1930), from S.A. in north Peru, Dept. Cajamarca in mtns., published in “Engl. Nat. Pflz.fam.” 1930, II: 18a:473, species name means “broad cloaked”; main stems about 10 cm tall, the rosette leaves small, fleshy, densely clustered, grayish-green, often red-tipped. Flowering stems are 25 cm racemes & have numerous small, rounded bract leaves. First placed in the old Series Racemosae & now in new Series 12 of same name; cytology: n=approx. 80.
=Cotyledon eurychlamis, (Diels, 1906), obs.
=*Ech. eurychlamis (Berger, 1930), current.

EXCELSA (Berger, 1930), in Peru in Dept. Ancash at 3600 M (about 18,000 ft.), name means “high or above” in reference to high alt. habitat (it grows in sight of Peru’s highest mountain, Mt. Huascaran, some 6,768 meters high); this species is in series 12 in 2003 Lexicon, but Pino cites this: “According to Walther E. excelsa belongs to series Elatae---”, but after new studies of the rediscovered plant this could be changed; cytology still not listed in 2003 thus: n=? (but the new plants will probably soon be tested to determine correct count). Main stems about 10 cm, medium rosettes, leaf color shiny medium green, inflorescences very tall spikes (one to 1.2 meters!); flower color scarlet with paler tips. Type- species by Weberbauer W 31489 B, but this species not in cultivation as late as 2003 (as no one could find it), but in 2006 it finally was relocated by Pino. For the latest details read the report by G. Pino written for Haseltonia #12 of 2006. He remarks that this is the largest Echeveria native to Peru, and he located it in nature for the first time since Weberbauer collected it in 1906. His new report gives many new details on this species, so should be the best authentic reference to it, and it will no doubt be introduced to cultivation at last due to this recent rediscovery. Its closest relative is Ech. andicola. Pino found the plants to have “usually single” rosettes, so it may not cluster, making propagation more difficult. Judging from photos this should make an excellent Ech. for cultivation by collectors.
=Cotyledon excelsa (Diels, 1906), obs. [in Engler’s Botan. Jahrbuch 37:412, 1906].
=*Ech. excelsa (Berger, 1930), current. [in Engl.& Prantl, Pflanzenfam, ed. 2,18a:473, 1930, & Poellnitz in Fedde Repert, Vol. 39, p.217,1936, & in Macbride, Flora of Peru, Vol. XIII, Part II, No. 3:1014, 1938] and in Haseltonia #12 with excellent photos by Pino.

expatriata (Rose, 1905), no natural habitat known, this is a hort. hybrid, (name means “out of country”, or “without a country”) probably a chance garden hybrid, first placed in old Walther series 1, #5, p.71, cytology: n=?, (counts not practical in hybrids)], pub. in N. Amer. flora, 1905, XXII: 26, see table #1 on p. 81 of Haseltonia #2, 1994 for Uhl’s latest test for this species. This is still widely found in nurseries, & collections. It is easy, and attractive, but not a wild species.
=Ech. expatriata [Rose, 1905], obs.
=*x Cremneria expatriata, Moran, comb. nov.[ in Baileya 1975 ], current. (Cremnophila linguifolia x E. amoena (microcalyx))

FIMBRIATA (Thompson, 1911), Morelos, Mex. in Sierra de Tepoxtlan, name means “fringed” (for leaves: see below), field collection number Trelease-674; was in old Walther series 2, sp.#50, page 186; but now in 2003 Lexicon it is listed in new Series 5 (Gibbiflorae), cytology: n=54. Species pub. in Transcriptions Academy of Science, St. Louis, 1911, XX:2:20. Main stems are unbranching 50 cm long, leaves green to brown-red, and minutely fringed (fimbricate); inflorescences cymose, 60 cm; flowers red to pink.
=*Ech. fimbriata [Thompson, 1911], current.

FULGENS (Lemaire, 1845), N. Cent. Mexico, widespread species: states Durango to Oaxaca, name means “gleaming”, first placed in old Walther series 2, #42, p. 159; now in new Series 5 (Gibbiflorae) in 2003 Lexicon, cytology varies a bit by variety (see below): n=27, 135, 162. ISI offering in CSSA Journ. Mar. '81, p.63 (ISI-1260), see Haseltonia #5, p.51, 1998. Two varities recognized in 2003 Lexicon (see below). Main stems 15 or more cm tall, rarely branching, leaves glaucous-green, usually heavily tinged with red or brown-red with margins often undulate and darker red; inflorescences cymose, 15 to 20 cm tall, flowers peach-red, petals with appendages near nectary (sometimes). Variety obtusifolia has smaller leaves, and these are mostly plain green.
=*Ech. fulgens (Lemaire, 1845), current.
=*Ech. fulgens var. fulgens (Lemaire, 1845), current for type species. (n=27, 162)
=Cotyledon retusa (Lindley, 1847; Baker, 1869), obs.
=Cotyledon fulgens (Lemaire, 1845; Baker, 1869), obs.
=Ech. retusa [Lindley, 1847], obs.
+*Ech. fulgens var. obtusifolia [Kimnach, 1998], current for the other variety. (n=27, 135, 162)
=Ech. obtusifolia [Rose, 1903], obs.
=Ech. scopulorum [Rose, 1905], obs.
=Ech. obtusifolia var. scopulorum [Poellnitz, 1936], obs.