ECHEVERIA LISTINGS L-M

By James E. Low

lagunensis (Munz, 19??). Walther saw that this was not an Echeveria and he republished it as follows:

=Ech. lagunensis (Munz, 1932), obs.

=Dudleya lagunensis (Walther, 1932), obs.

=Dudleya arizonica (Rose, 1923), obs.

=*Dudleya pulverulenta ssp. arizonica (Moran, 1945), current.

Ech. “lanceolata var. incepta” (Jepson, 1936), but not an Echeveria.

=*Dudleya cymosa (see Madrono of Oct. 1987, p. 335), current.

LAUI (Moran & Meyran, 1976), SW Mex., state of Oaxaca, pub. in Mex. C&S Journ. 1976, 2:57, photo fig. 4, p. 259, CSSA Journ., Nov. 1988, color ph.: CSSA Journ. Mar. 1995, p.87 with text; ISI offering with color ph. in CSSA Journ., p.101, Mar. 1980 (ISI 11-99), name honors Alfred Lau who discovered it in almost inaccessible, dry, hot low altitude (500 m) canyons full of thorn bushes; first placed in Series Angulatae, but now in new Series 11 (Pruinosae) in 2003 Lexicon; cytology: n=15, see Haseltonia #6, page 65. Plants usually un-branched, leaves white glaucous to pink glaucous, dead leaves persist on stem; the inflorescences are multiple branched, short cincinni, flowers red and pruinose (frosted). This species is considered to be one of the most beautiful in the entire genus. In cultivation the plants are easily damaged, and require careful culture. Usually no stem offsets for use in propagation, but some say that new plants are easy from detached leaves, but some say it is not easy, and requires special handling.

=*Ech. laui [Moran & Meyran, 1976], current.

 

LEUCOTRICHA (Purpus, 1914), E. Cent. Mex. state of Puebla, (name means “white haired”), this is one of the hairy species similar to Ech. pulvinata, but with shorter leaves and different chromosome number; pub. in Monatsschrift Kakteenkunde (German), 1914, 24:65; see also in Amer. Journ. Bot. 1992 on p. 56 Uhl lists this sp. (1976 & 82) and gives the cytology as: n=38 (an “auto-tetraploid”), first listed in old series 13 (Racemosae), but now in new Series 4 (Echeveria) in the new 2003 Lexicon. The bare stems branch above ground to lax rosettes of a few fuzzy leaves. Leaves green and are dense with white “hair”; the infl. is sub-spicate to sub-paniculate to 40 cm tall, flowers red.

=*Ech. leucotricha [Purpus, 1914], current.

 

 

LILACINA (Kimnach & Moran, 1980), NE Mex. state of NL, (means “lilac colored” for leaves), pub. in CSSA Journ. Jul. '80, p.76, see also CSSA Sep. '81 (BW photo), & see CSSA Jul. 1994, p. 172, in Haseltonia #4 of 1996 Uhl discusses this sp. and lists it in Series Urceolatae, n=27 (diploid), but Kimnach moved it into his new series 16 (Urbiniae) in 2003 Lexicon. It has attractive rosettes, not branching; although it was new it soon became popular in nurseries everywhere. The base leaf color is difficult to see through the white to pinkish powder, but it is said to be “brownish-olive-green” when powder is removed; inflorescences are single to double cincinni, flowers pink. Color photo on p. 199 of CSSA Journ. of Sept. 2007.

=*Ech. lilacina [Kimnach & Moran, 1980], current.

 

 

lindsayana (Walther, 1972), from Mexico, possibly in state of Jalisco; first listed in old Walther series 2, #11, p. 90, pub. in book ‘Ech.’, 1972, p.90; see CSSA Journ. Mar, 1980, p.58 (name honors Geo. Lindsay, see p.58, CSSA Journ. Mar. 1980), now listed in new Series 16 (Urbiniae) in 2003 Lexicon. See Haseltonia #4 of 1996 on p. 79, cytology is: n=27. This sp. now combined under a different species name.

=Ech. lindsayana [Walther, 1972], obs.

=*Ech. colorata var.colorata (Kimnach,1980). Current. See full data under this name.

linguaefolia (Lemaire, 1863), Now placed in a different genus: linguifolia (Moran, 1975), this species name means “tongue leafed”. For details read CSSA Journ. May, 1978, p.139-147. From E. Cent. Mex. state of Mex. at Barranca de Mexicapa, n=33, first placed in old Walther series 1 (Paniculatae), #1 later moved to Induplicatae; Moran moved this sp. to new genus Cremnophila on p.139 of CSSA Journ. of May, 1978.

=Ech. linguaefolia [Lemaire, 1863], obs.

=Cremnophila linguifolia [Moran, 1975], current.

LONGIFLORA (Walther, 1959), SW Mex., state of Guerrero (?): this species was published by Walther based on a cultivated plant thought to have come from Guerrero, species name means “long flowered”, first placed in old Walther series 2, #52, p.192, it is now listed in the new Series 5 (Gibbiflorae) in 2003 Lexicon. Species published in CSSA Journ. 1959, 31:101; cytology: n=27, plants usually unbranched, leaves green, tinged violet, and more or less glaucous; inflorences rather tall (to 75 cm) and cymose, 3 branched; the odd long flowers are purplish below to rose above and glaucous. Kimnach had the following to say in 2003 Lexicon about this species: “Distinct in the E. gibbiflora alliance because of its long narrow corolla lacking in a yellow or orange coloration. Similar flowers are found in E. scheeri, which otherwise seems distinct”.

=*Ech. longiflora [Walther, 1959], current.

longipes (Walther, 1935), E. Cent. Mex., state of Hidalgo, means “long footed”, pub. in CSSA Journ. 1935, 7:36, first placed in old Series Mucronatae. This species was combined into another species in 2003 Lexicon by Kimnach.

=Ech. longipes [Walther, 1935], obs.

=*Ech. paniculata var. maculata [Gray, 1852; Kimnach, 2003], current.

LONGISSIMA (Walther, 1938), E. Cent. Mex., S. Puebla state; pub. in CSSA Journ. 1938, 9:147; name means “longest” in ref. to the long style (inside flower), this species comes in two subspecies as listed below. See ISI offering in CSSA Journ. 1976, p. 90, (ISI-965). First placed in old Series Longistylae, now in new Series 6 of same name in 2003 Lexicon; cytology: n=42, plants rarely branching, leaves dark green often with red margins, medium inflorescences with one to 3 cincinni, flowers large, orange with green tips. Flowers continue to become larger with age and are largest of entire genus. This is considered to be a choice looking plant for collectors, but difficult, slow growing, and hard to propagate.

=Ech. harmsii var. multiflora (Walther, 1935), obs.

=*Ech. longissima var. longissima (Walther, 1938), current.

+*Ech. longissima var. aztatlensis (Meyran, 1982), of Sierra Mixteca, Oaxaca state of SW Mex. pub in Mexican C&S Journ, 1982, 27:33. This ssp. was found by Otero as his field number 25. Differs from type species in having smaller leaves, and plant is caespitose, & old flowers resemble Chinese lanterns. This ssp. is easier to propagate and easier to grow. Current.

 

 

LOZANOI (Rose, 1905), W. Cent. Mex., Jalisco state, in mts., originally placed in old Walther series 2, sp. #37, p.151, pub. in North American Flora, 1905, XXII:23, (name honors F. Lozano who found it with Pringle in 1903); species never seen again after first discovered probably due to goats, thus it cannot be tested so: n=? No plants still in cultivation, but some description was made so we know it had a low rosette & at least the central leaves were copper colored; inflorescences 30 to 45 cm tall in a cyme with 3 to 7 branches, sepals unequal, flowers copper colored also. All attempts to find it again in original habitat and nearby have failed, but enough hope of finding it remains to continue listing it in the latest Lexicon. Old spelling was “lozani” which is incorrect Latin, so the spelling above is proper.

*Ech. lozanoi [Rose, 1905] current.

lurida (Haworth, 1831), E. Cent. Mex. near Veracruz, (name means “pale”), pub. in Taylor's Phil. Mag. 1831, X:418. In 2003 Lexicon reduced to synonymy with Ech. racemosa var. racemosa.

=Cotyledon lurida [Haworth], Baker (1869) obs.

= Ech. lucida [Steudel, 1840], obs.

=*Ech. racemosa var. racemosa, current.

LUTEA (Rose, 1911), NE Mex. state of SLP & Zac. in Sierra de Alvarez, (name refers to its saffron yellow flowers), pub. in Journ. Wash. Acad. Sci., 1911, I: 1.2:268; now in the new Series 1 (Angulatae) in the 2003 Lexicon; cytology: n=12. Rarely branching, densely rosulate, leaves green to brownish; inflorescences 1 or 2 cincinni, 28 to 75 cm tall, flowers yellow. This species attractive, but difficult as it is from hot and dry areas. Flowers said to be especially attractive.

=*Ech. lutea [Rose, 1911], current.

+var. fuscata [Walther, 1972], obs.; was retracted in 2003 Lexicon.

 

LYONSII (Kimnach, 2006), This is a species nova published in CSSA Journal of Sept. 07 on page 215 to 219 with plenty of data and photos. It is named to honor the co-discoverer Gary Lyons. Kimnach said the species has been known since discovery in 1971, but it was not published until now. Series Angulatae, n=12, related to E. shaviana & E. bifida, leaves very pale, glaucous, offsetting, easy in culture, flowers red-scarlet to pinkish on 8 inch stems, infl. has several cincinni that arch over pendent, habitat: Tamaulipas state in NE Mexico, between Tula and C. Victoria. A natural hybrid is known, but not named, and is somewhat similar but larger, and leaves more purplish, HNT-41537 =Uhl-2845 from near same habitat. Note that good color photos are given of both types in the publication article in the CSSA Journal: the type species is shown on p. 217, and the hybrid on p. 219. Here the two look very different, indeed.

=*Ech. lyonsii [Kimnach, 2006], current.

MACDOUGALLII (Walther, 1958), SW Mex. state of Oaxaca, pub. in CSSA Journ. 1958, 30(3):87, name honors botanist Tom Macdougall. ISI in CSSA Journ. Mar.1995 (ISI 95-27). This is an odd species that does not look much like an Echeveria, so it was placed first in another genus. It is now firmly in Echeveria, and the 2003 Lexicon places it in the new Series 8 (Nudae), and the cytology of plants from over its native range is complex: n=19, 34, 40, 42, and 50 have all been found! Plants are much branched, weak rosettes; inflorescences are short racemes, flower color is given as “reddish-yellow”.

=Ech. sedoides [Walther, 1972], obs.

=*Ech. macdougallii [Walther, 1958], current.

 

MACRANTHA (Standley, & Steyermark, 1944), Guatemala, Dept. Jalapa at 2000 m alt., (name means “large flowered”), pub. in Field Museum notices in Bot. Ser, 1944, 23:4:159; plants shrubby, but little branching, leaves a bit hairy and yellow-green with rose margins, stout inflorescences to 45 cm tall but only 3 flowers, flower color not listed, but petals are called hirsute. This is still a little known species needing more study. Walther thought it could be the same as Ech. pringlei. It appears that this species has not been found in recent times, and it is not in cultivation.

=*Ech. macrantha [Standley & Steyermark, 1944], current.

 

maculata (Rose, 1903), said to be widespread in Centr. Mex., (name means “spotted”), pub. in Bulletin NY Bot. Gard., 1903. III:7. This sp. now combined into another species; see Haseltonia #5:51, 1998.

=Ech. maculata [Rose, 1903], obs.

=*Ech. paniculata var. maculata [Gray, 1852; Kimnach, 1998], current.

MAXONII (Rose, 1909), Guatemala in Dept. Quetzaltenango at 2200-3300 m alt., & also found in Honduras; publ. in Contr. US Nat. Herbarium, 1909, XII:395. Like some other Central American species this has a long stem below the leaves which are barely rosetted, main stem is little branching; leaves green and minutely papillose, inflorescences are a raceme or are subpaniculate to 60 cm tall with a few scarlet flowers, cytology: n=26; this species now listed in the new Series 8 (Nudae) in the 2003 Lexicon.

=Cotyledon acutifolia non Lindley, [Hemsley, 1880], obs.

=*Ech. maxonii [Rose, 1909], current.

 

 

MEGACALYX (Walther, 1959), SW Mexico in state of Oaxaca at Teotitlan del Rio at 1940 m elev., name means “huge calyx”, pub. in CSJA Journ. 1959, 31:50-51; now in the new Series 12 (Racemosae) in the new 2003 Lexicon, cytology: n=20. Rarely branching, leaves green to slightly glaucous-blue, the inflorescences a raceme to 45 cm tall, flowers a dull greenish-yellow. This species said to be difficult due partly to mealy bug infestations.

=*Ech. megacalyx [Walther, 1959], current.

meyraniana (Walther, 1959), E. Cent. Mexico in E. Puebla state on limestone hill, pub. in Mex. C&S Jour. 1959, IV:29, first placed in old Walther series 2, sp. #29, p. 134, honors Mex. C&S botanist J. Meyran. Note: in Haseltonia #3 of 1995 on p. 37 Uhl combined this as a ssp. of Ech. subalpina, and the 2003 Lexicon drops it even as a subspecies.

=Ech. meyraniana [Walther, 1959], obs.

=*Ech. subalpina [Rose & Purpus, 1910], current.

microcalyx (Britton & Rose, 1911), E. Cent. Mex., Vera Cruz to Puebla states, name means “tiny calyx”, first placed in Walther series 1, #2, p.66; pub. in Contr. US Nat. Herbarium, 1911, XIII:295. Prof. Uhl thought that this sp. should be combined with Ech. amoena, but did not do it himself: see in Haseltonia #2 if 1994 on p. 85. These and the newer Ech. prolifica are closely related, and all are related to Moran’s new genus Cremnophila (more studies are needed). Uhl handles this only as the same sp. as Ech. amoena, Haseltonia #2, old series 1 (Paniculatae), collected plants had n=33, 66. In the new 2003 Lexicon this species was indeed combined into Ech. amoena, and then placed in the new series 10 also called Paniculatae. See that listing for details.

=Ech. microcalyx [B & R, 1911], obs.

=*Ech. amoena [De Smet, 1875], current.

MINIMA (Meyran, 1968), E. Cent. Mexico in Hidalgo state, name means “least” (in size), pub. in Mexican C&S Journ. 1968, 13: 47-62; see CSSA Journ. Jan. 1981, BW photo & caption text; CSSA Journ. Mar. 1977, ISI-1014 (text only); and Haseltonia #3 of 1995 on p. 35, cytology: n=28 (tetraploid). A freely branching species with small, dense and attractive rosettes; leaves glaucous green with red accents, inflorescences short cincinni, flower color rose at base to yellow above. This is now a popular species in cultivation.

=*Ech. minima [Meyran, 1968], current.

 

minutiflora (B&R, 1903), for an odd plant that was in time found to be in incorrect genus.

=Ech. minutiflora [Britton & Rose], obs.

= Ech. tepacensis (Poellnitz, 19??). obs.

=*Thompsonella minutiflora (Rose, 1909), current.

 

MONDRAGONIANA (Reyes & Brachet, 2009) published in Cactáceas y Suculentas Mexicanas, 54(3): 86, ills., 2009. Found in Miahuatlan in the mountains of the southern part of Oaxaca, Mexico, growing in protected places like rocky canyons in pine and spruce forest at 3000 m and higher up.
This is a glabrous plant with a short stem, rosette 6 – 14  cm in diameter, leaves ca 5 x 3 cm, obovate, cuspidate, inflorescences 20 – 30 cm high, racemose, pedicels up to 18 mm long, sepals unequal, corolla ca 14 mm long, yellow. Superficially the rosettes resemble some forms of E. secunda and there is also some resemblance to E. sessiliflora but E. mondragoniana is clearly distinguished from both species by the type of inflorescence and its yellow flowers on long pedicels.
It is provisionally placed in Series Racemosae.
=*Ech. mondragoniana [Reyes & Brachet, 2009] current.

 

MONTANA (Rose, 1903), SW Mexico at higher elev. in states of Chiapas & Oaxaca & into Guatemala, pub. in Bulletin NY Bot. Gard., 1903, III:6, the Latin name refers to its being a “mountain type” plant, see Haseltonia #3 of 1995 on p. 28 where Uhl lists this as having n=22 (same as in new Lexicon), listed in new Series 8 (Nudae), plants on long, branching stems (shrubby) as much as 50 cm tall to rosettes which are loose with smooth, green leaves, the infl. a spike to 60 cm tall above leaves, flowers reddish with yellow margins.

=Ech. nuda var. montana [Poellnitz, 1936], obs.

=*Ech. montana [Rose, 1903], current.

MORANII (Walther, 1972), SW Mex. Oaxaca state, near Totolapan, 1150 m elev., (name honors Prof. Reid Moran, USA Botanist), pub. in Ech. 1972, p. 347; first placed in old Series 11, now new series 12 (Racemosae). Cytology: n=24; plants usually not branching, leaves grey-green with red margins & keels & with scattered red dots, the inflorescences are almost horizontal, 15 to 30 cm long, and the flowers pendent on that, flowers red to scarlet.

=Ech. proxima [Walther, 1972], obs.

=*Ech. moranii [Walther, 1972], current.

Mostel’s species nova, 1996; BLM-0834, (unnamed and unpub.) of Burl Mostel, owner of Rare Plant Research of Portland. Ore. offers this in his 1996 catalog. He said it resembles Graptopetalum paraguayense. Thus, it may indeed be a Graptopetalum and not an Ech.! Current status uncertain.

MUCRONATA (Schlechtendal, 1839), widespread E. Cent. to N. Cent. Mexico (in ten of the states!), especially in state of Hidalgo; name means “pointed” (for leaf tips); see CSSA Journ. Nov. 1993, p. 272; published in Linnaea, 1839, XIII:411, see CSSA Journ. Nov. 1993, p. 272. Uhl thinks it may in time be combined with Ech. crassicaulis and Ech . platyphylla; this species was placed in old series Mucronatae & in new Series 7 (of same name) in 2003 Lexicon, Cytology: n=16 or 32; roots thick, plants rarely branching, all leaves may fall entirely at times, leaves green and a bit glaucous, often spotted red-brown, inflorescences grow as long spikes to 50 cm.; flower color amber to light yellow. Note that this species is variable, and some from farther north with small flowers are under consideration as a possible new species.

=Cotyledon mucronata [1839; Baker 1869], obs.

=Ech. crassicaulis [Walther, 1935], obs.

=*Ech. mucronata [Schlechtendal, 1839], current.

MULTICAULIS (Rose, 1905), SW Mex. state of Guerrero, means “multi-stemmed”, pub. in Contributions US Nat. Herbarium, 1905, VIII:294; now in new Series 8 (Nudae) in 2003 Lexicon, chromosome count of n=21 was by Uhl in 1982, and holds true today. This is a freely branching species on elongated and rough main stem, rosettes loose and poorly formed, leaves shiny, dark green with red margins and tips, inflorescences are either a raceme, a spike or even a thyrses to 25 cm long; flowers carmine to scarlet.

=*Ech. multicaulis [Rose, 1905], current.

 

multicolor (Uhl, 1992), Venezuela near Merida at 3080 m elev., pub. in CSSA Journ. May, 1992, p. 120-124; see also the article by Uhl in CSSA Journ. Mar. 1993, p.81 to 87 (Form 4) with text & photos related to this species, (name refers to “many colored” leaves and stems), this species was closely allied to the Ech. bicolor complex. Multicolor was in old series Racemosae, moved to new Series 8 (Nudae); Cytology: n=21 (diploid), or rarely n=42, (tetraploid). Note: Kimnach discarded the name of Ech. multicolor entirely in the 2003 Lexicon for (?) reasons, and he reinstated an older name that he at one time had said was invalid.

=Ech. multicolor [Uhl, 1992], obs.

=*Ech. recurvata [Carruthers, 1974], current.