ECHEVERIA LISTINGS P

By James E. Low

pachanoi (Rose ex Poellnitz 1935); little original data available but probably of Ecuador, fully published by Fedde in Berlin after Rose’s death (see page 187 of the Fedde Report 1938). Name honors Abalardo Pachanoi, plant collector of Ecuador; now combined into another species in 2003 Lexicon by Kimnach.

=Ech. pachanoi, [Fedde, 1938], obs.

=Ech. quitensis var. quitensis [Lindley, 1852], current.

 

pachyphytum (Morren, 1874). East central Mexico, see CSSA Journ. May 1991, p. 119.

=Ech. pachyphytum (Morren, 1874), obs.

=*Pachyphytum bracteosum [Klotzsch, 1841], current.

PALLIDA (Walther, 1938), probably from southern Mexico (in Chiapas state ?) exact location not known and now in controversy, one field number given is FO-144. Species first placed in old Walther series 2 (Secundae), sp. #53, p. 194; now in new Series 5 (Gibbiflorae) in 2003 Lexicon; cytology: n=54. Species published in CSSA Journ. 1938, 10:14. Latin name means “pale in color” referring to leaves. This is a large species and when well grown and in full flower it can be one of the most beautiful plants in genus Echeveria for the huge clusters of lovely pink flowers. Blooms in winter. Today this species is still little known in collections. Old plants can become “leggy” and not attractive, so they need good management; but offsets are borne freely. Main stems to 10 cm or more long and 1 ½ cm in dia.; rosettes up to 20 cm in dia.; inflorescences cymose-paniculate; flowers pink to red. Good plants for cultivation should be carefully selected for color, pink usually preferred.

=*Ech. pallida ([Walther, 1938], current.

 

 

 

palmeri (Rose, 1903), NE Mexico in state of SLP at 2100 m elev., old Walther series 2, sec. Gibb., sp. #47, pub. on p.179, pub. in Bulletin NY Bot. Gard., 1903, III:10, see CSSA Journ. Nov. 1992, p. 303 (BW photo p. 312), photo CSSA Journ. p. 243, Sept. 1997: now considered merely a green form of the usually frosted leaved E. subrigida, thus name E. palmeri is no longer to be used. For details read in the Sept, 1997 issue of the CSJA pages 239 to 244. The older names were:

=Ech. rosei (not rosea), [Nelson & Macbride, 1913], obs.

=Ech. palmeri [Rose, 1903], obs.

=Ech. angusta [Poellnitz, 1936], obs.

=Ech. subrigida fma. palmeri [Glass, 1997], probably still true for green leaf form.

=*Ech. subrigida [(Robinson & Seaton) Rose 1903], current.

PANICULATA (Gray, 1852), N. Cent. Mexico in states of Chihuahua, Durango, & Zacatecas; its name means “panicalled” (in ref. to flower cluster); species published in Plant. Wright., 1852, 1: 76, first placed in Walther’s old series #1, but moved to new Series 15 (Thrysiflorae) in 2003 Lexicon; cytology: n=28, roots fusiform & fleshy, but little true stem; leaves thin & green, inflorescence of type-species consists of 1 to 4 cincinni to 15 cm tall, flowers called glaucous pink; but var. maculata differs from type sp. in its more succ. leaves, more expanded sepals, and pure yellow fl. and the inflorescences are in thryses form with this being either subpaniculate below and spicate-racemose above, or having mostly 2 flowered branchlets according to Kimnach.

=*Ech. paniculata [Gray, 1859], current.

=*Ech. paniculata var. paniculata [Kimnach, 1998], current for type-species only.

=Cotyledon grayi [Baker, 1869], obs.

=Ech. grayi [Morren, 1874], obs.

=*Ech. paniculata var. maculata [Gray, 1852; Kimnach, 1998], current for this var. only.

=Ech. maculata [Rose, 1903], obs.

=Ech. longipes [Walther, 1935], obs.

 

 

PAPILLOSA (Kimnach & Uhl, 1983), Guatemala, SE of San Cristobal at over 3000 m elev., name ref. to fuzzy leaf & stem surfaces, pub. in CSSA Journ. Jan. 1983, p. 27, in the 2003 Lexicon this species is listed in new Series 12 (Racemosae); cytology: n=15, but Kimnach remarks: “a species without close allies”, so it must seem odd to him. It is a branching type with proliferous rosettes above 5 inch stems, the leaves are papillose with odd semiglobose hairs, leaves green with somewhat red margins; inflorescences 2 sided racemes; even the red-orange flowers are papillose.

=*Ech. papillosa [Kimnach & Uhl, 1983], current.

parrasensis, (Walther, 1959), E. Cent. Mexico in state of Coahuila; first listed in old Walther series 2 (Secundae), sp. #14, p. 93 (fig. 34, p. 93), means "of Parras" (a place in Coahuila state).

=Ech. parrasensis [Walther, 1959], obs.

=*Ech. cuspidata not of Rose but of [Purpus, 1907], current.

PEACOCKII, (Croucher, 1874), N. Cent. Mexico in state of Coahuila; pub. in Garden Chron. 1874, I:674; see Haseltonia #3, p.28 & 47,Walther placed this sp. in his series Pruinosae, now in new Series 11 of same name in 2003 Lexicon; cytology: n=15. Name honors J. T. Peacock, succulent plant collector of England. For BW photo refer to p. 168 of CSSA Journal on Nov. 1964, fig.2. Plants rarely branching, leaves densely rosulate, bluish-white frosted with red margins, inflorescences with 1 to 4 cincinni to 15 cm tall, flowers glaucous pink. This species grows in hot, dry areas mostly in shade.

=Cotyledon peacockii [Baker, 1874], obs.

=Ech. desmetiana [Morren, 1874], obs.

=Cotyledon desmetiana [Hemsley, 1880], obs.

=Ech. subsessilis [Rose, 1905], obs.

=*Ech. peacockii [Croucher, 1874], current.

 

 

PENDULIFLORA (Walther, 1958), S. Mexico in state of Oaxaca at Cerro Gavilan at 5000 ft., name means “hanging flower”, pub. in CSSA Journ. 1958, 30(5):151; MacDougall field number B-174; Series not mentioned at publication, but now listed in new Series 12 (Racemosae) in 2003 Lexicon; cytology: n=22 (see AJB of 1982, p. 846.), and same in Lexicon. Plants shrubby to 30 cm tall, no hairs, leaves scattered to subrosulate, green; infl. racemes to 30 cm tall, flowers pink to red.

=*Ech. penduliflora [Walther, 1958], current.

PENDULOSA (Kimnach, & Uhl, 1992), Venezuela in Dept. Merida 30 km west of Merida the city, name means “hanging” (plant): stems up to 2 meters (8 ft.) always hanging downward; pub. in CSSA Journ. Jul. 1992, p. 200; ISI offering Mar. '93. p. 74, ISI #93-31; in 2003 Lexicon it is listed in new Series 8 (Nudae); cytology: n=42, related to E. bicolor, but differs in hanging habit, erect sepals, and in chromosome count; stems begin by much branching, then growing downward to as much as 2 meters or more with dense aerial roots, leaves few and these only near stem tips, bluish-green glaucous, margins reddish; the inflorescences are upcurving racemes to horizontal on pendent stems; flowers yellowish-peach below, pinkish-yellow near tips.

=*Ech. pendulosa [Kimnach & Uhl, 1992], current.

 

PEREZCALIXII (Jimeno-Sevilla & P. Carrillo, 2010) published in Brittonia, 62(4), pp. 303-308, 2010. Found in the southern part of the Sierra Madre Occidental in northern Jalisco and in southern Nayarit and Zacatecas, growing in tropical deciduous forest on volcanic rocks in poor soil at 1500 – 2000 m alt.
This is a glabrous and pruinose plant, stem only 1,5 cm tall, leaves obovate-spatulate, with hyaline margins, inflorescences paniculate, up to 43 cm high with normally 3 cincinni with up to 9 flowers, pedicels up to 14 mm long, sepals unequal, rather thick, pruinose, corolla slightly urceolate, ca 12 mm long, orange-pink to pink, pruinose, petals inside yellow.
Placed in Series Gibbiflorae and compared with E. fulgens, from which it differs by its acaulescent habit, pruinose leaves with hyaline margins and longer pedicels and sepals.
=*Ech. perezcalixii [Jimeno-Sevilla & P. Carrillo, 2010] current.

 

PERUVIANA (Meyen, 1834), SE Peru, Dept. Tacna near Palca on borders Bolivia & NW Argent., variable sp.: many forms known from this general area, all still unnamed. This species rarely offsets (unlike the nearby Ech. decumbens). Peruviana is in a group allied to Ech. eurychlamis & excelsa. Species was pub. in Reise um die Erde 1:488, in adnot., 1834 (the Latin name honors Peru); now listed in 2003 Lexicon in new Series Racemosae, with probable cytology of: n=approx. 50, leaf color uncertain (old data incomplete), as to inflorescences it is known only that there were leaf-like bracts in it, flower color given as purple. This species has not been found near the stated type-locality in recent times (may be extinct there), but similar plants are found in the general region, yet they differ in several details. For example, the plant Walther used to describe for his 1972 book was in reality from Argentina! Thus, old data are not now accepted. More field work is needed. The closest plant to original site found recently seems to be field number: Hutchinson 7176 from 62 km NE of Tacna. We know it has green leaves with red edges, red petals, and the count of n=approx. 50 (data shown above was from it). Similar plants from Argentina can vary in details, but typically most have green leaves, sometimes glaucous, but flowers may be anything from yellow to orange, or even red.

=Cotyledon peruviana [Baker 1869], obs.

=*Ech. peruviana [Meyen, 1834], current.

PILOSA (Purpus, 1917), E. Cent. Mexico in state of Puebla in the Sierra de Mixteca in mts.; name means “hairy”; pub. in Monatsschr. Kakteenkunde 27, 1917, p.146, first in old series Ciliatae, now in 2003 Lexicon in new Series 3 of same name; n=24, see in Haseltonia #5, p.22 & 29. This is another of the hairy types: hirsute everywhere except inside flowers, plants rarely branch, rosettes large but lax, green under hairs; the inflorescences cymose, about 5 branches each with few flowers; flowers scarlet outside, shading to orange, and yellow at tips. Similar to E. coccinea but has shorter stems below and type of inflorescence called “cymose-paniculate”.

=*Ech. pilosa [Purpus, 1917], current.

 

PINETORUM (Rose, 1905), S. Mexico in states of Chiapas & Oaxaca & into Guatemala, the Latin species name means “pine cone” (reason for this name not clear); pub. in  N. Amer. Flora, 1905, 22(1):20, 1905. Fleshy root, little main stem; little branching; leaves yellow-green often falling in dry season; inflorescence a spike to 30 cm, flowers scarlet at base to yellow above, cytology: n=14. This species is difficult to grow. It is closely related to E. sessiliflora, (but it has non fleshy roots, and blueish, glaucous leaves & its n=15).

=Ech. sessiliflora, var. pinetorum [Poellnitz, 1936], obs.

=Ech. huehueteca [Standley & Steyermark, 1944], obs.

=*Ech. pinetorum [Rose, 1905], current.

PITTIERI (Rose, 1911), Cent. America in Guatemala, Nicaragua, & Costa Rica; often grows epiphytic on trees in dense forests; species published in Contributions US Nat. Herbarium, 1911, XIII:296, in nature it grows with Ech. chiapensis (now synonym of E. rosea) & Ech. rosea. Name honors H. F. Pittier who collected it first (as his P-1880). Main stems branched, and up to 5 inches tall, leaves in loose rosettes, green often tinged brown; the inflorescences a 2-sided spike with many red flowers. Cytology: n=62.

=*Ech. pittieri [Rose, 1911], current.

 

PLATYPHYLLA (Rose, 1903), E. Cent. Mexico in State of Mexico (Fed. Dist.) to state of Hidalgo; means “flat plant”, pub. in Bulletin NY Bot. Gard. 1903, III:7, Prof. Uhl thinks this and Ech. crassicaulis and Ech. mucronata may eventually all be combined as a single spcies, but in 2003 Lexicon Kimnach does not seem to agree as he compares it to E. paniculata, saying platyphylla differs in having smaller leaves of differing color, and in type of infl. First placed in series Mucronatae, but now in new Series 15 (Thyrsiflorae) in 2003 Lexicon; Cytology: n=28. Root fleshy with little main stem, plants rarely branch, leaves often lacking in dry season, color grey-green; inflorescences are long spikes, with about 15 crowded flowers of yellow color turning more reddish in time.

=*Ech. platyphylla [Rose, 1903], current.

 

planiflora (Berger, 1930), SW Mexico; name means “flat flower”. This is not an Echeveria.

=Ech. planiflora [Berger, 1930], obs.

=*Thompsonella platyphylla [Rose, 1909], current.

potosina (Walther, 1935), NE. Mexico, named for state of San Luis Potosi & found near the city of that name; pub. in CSSA Journ. 1935, VII: 61, Walther sp. #16, p. 104 (fig. 43, p. 105), see Haseltonia #4 of 1996 on p. 80, was in old series Secundae, later moved into Uhl’s revised Series Urceolatae, cytology: n=60-62 for one plant & n= about 115 & for another and n= about 117 for still another plant n =112 (see in AJB of  1982, p. 846). In the 2003 Lexicon this species, E. potosina, is listed as being combined into another species as shown below.

=Ech. potosina [Walther, 1935], obs.

=*Ech. elegans [Rose, 1905], current.

PRINGLEI (Rose, 1903), NE Mex. to N. Cent. to W. Cent. Mexico in states of Jalisco, SLP, Durango in mts. at 4500 m alt., honors early plant explorer Cyrus Pringle; species pub. in Bulletin NY Bot. Gard. 1903, III:6, p.561, see in Amer. Journ. Bot. 1992, now listed in new Series 4 (Echeveria) in 2003 Lexicon; this species comes in 3 varities as listed below; cytology for all: n=46 (auto-tetraploid). The first (var. longisepala) is from Jalisco state in W. Cent. Mexico at 1800 m and is tall and has smooth leaves and longer sepals (as its name implies), main stem is long and thick, leaves green and glabrous (hairless!), flowers orange-red; and the 2nd  (var. parva) is from Durango state of N. Cent. Mex. at 1350 m and has very small, reddish, also glabrous (hairless!) leaves [the Latin parva means small or dwarf], flowers light orange; while the type-species (var. pringlei) is from Jalisco state and is not hairless, but minutely pubescent, flowers orange-red to scarlet. Note: Kimnach says that “Ech. macrantha” may be a synonym for Ech pringlei, thus he leaves the matter unsettled at this time. Later decisions may combine these into a single species. In any case in nature these species are variable, sometimes difficult to separate.

=Cotyledon pringlei [Watson, 1890], obs.

=*Ech. pringlei, [Rose, 1903], current.

=*var. pringlei [Rose, 1903], valid for type-species only.

+*var. longisepala (Kimnach, 1998), var. nova in Haseltonia #5, p. 51, valid for new large form.

+*var. parva (Kimnach, 1998), current for small var. only (var. nova also in Has. #5, p.51).

 

 

 

PROCERA (Moran, 1967), SW Mexico in Oaxaca state; means “tall growing”, [probably allied to genus Pachyphytum, according to Moran due to “appendage” on each side of the nectary. Kimnach seems to doubt this because, as he points out 2 other species have this feature and are not closely related]. Published in CSSA Journ. 1967, 39(5), p.182. Plants hairless, cytology: n=22, listed in new Series 8 (Nudae) in 2003 Lexicon; lower stems thick & up to 2 meters tall! Large rosettes up to 30 cm in diameter, leaves plain green, inflorescences subspicate and to as much as 1 ½ meters tall, conspicuous bract leaves between flowers, the flowers red.

=*Ech. procera [Moran, 1967], current.

PROLIFICA (Moran & Meyran, 1978), E. Cent. Mex. Discovered in cultivation in Mexico, not yet found in the wild. Published in CSSA Journ Nov. 1978, p. 289; name refers to its heavy clustering habit, it is a newer member of the same group as Ech. amoena, and microcalyx (now synonym of E. amoena). Uhl thinks those 2 should be combined, but prolifica is just different enough to be a valid species (Chromosome study shows all 3 to be related to Moran’s new 2 sp. genus Cremnophila). Read p. 84 to 86 of Haseltonia #2 if 1994, first listed in old series 1 (Paniculatae), and now in new Series 10 of same name in 2003 Lexicon, Cytology: n=33. The bract leaves easily fall and start new plants. The mature plants send out stolons to grow many plants as sprouts around mother rosettes, leaf color a light, glaucous green, inflorescences are decumbent, cymose and 15 to 25 cm tall, dense cymes to 3 cm wide, each with 2 to 3 cincinni, and each of these with up to 4 unique bell-shaped flowers of yellow color.

=*Ech. prolifica [Moran & Meyran, 1978], current.

 

proxima (Walther, 1972), now combined into another species:

=Ech. proxima, [Walther, 1972], obs.

=*Ech. moranii [Walther, 1972], current.

PRUNINA (Kimnach & Moran, 1981), S. Mexico in state of Chiapas; name means “frosted” (for white leaves), pub. CSSA. Journ. Nov. 1981. p.294; first listed in old series Retusa, but now in new Series 5 (Gibbiflorae) in 2003 Lexicon, cytology: n=28. Lower stem thick and up to 5 inches tall, medium sized rosettes of heavily violet glaucous young leaves, older outer leaves loosing glaucous coating showing reddish-green color; the inflorescences are cymes with just 1 or 2 cincinni, medium tall and glaucous also, large bract leaves between flowers, pink flowers; cytology: n=28+1. This species is most closely allied to E. scheeri & to E. fulgens. Color photo on p. 198 of CSSA Journ. of Sept. 2007.

=*Ech. prunina [Kimnach & Moran, 1981], current.

pubescens (Schlechtendal, 1839), E. Cent. Mexico in States of Puebla & Hidalgo; means “downy or fuzzy” (leaves), pub. in Linnea, 1839, XIII:411, Series 13, n=23 by Uhl in 1956 & n=25 by Uhl and Moran in 1976. Note: in the 2003 Lexicon Kimnach writes in regard to this species as follows: “E. pubescens, supposedly with wider leaves, cannot be maintained, because of intergrading forms. Especially wide-leaf forms are found along the Ixtlan-Oaxaca highway and a white-haired form is found near San Luis Atolotitlan, Puebla”. He then combined E. pubescens into an even older species:

=Ech. pubescens [Schl., 1839], obs.

=*Ech. coccinea [De Candolle, 1828], current.

Ech. x pulchella (Berger, 1904), this “species” was found to be a “hort.-type”, described from cultivation of plant from Italy, not found in nature, (name means “beautiful”), Walther placed it in his old series 1, sp. #4, p. 70, pub. In Gartenflora, 1904, 53:206. In the 2003 Lexicon, after more study Kimnach had this to say: “Similar to x Cremneria expatria and, like it, a probable hybrid with E. amoena or Cremnophila linguifolia as one parent, and thus perhaps belonging to x Cremneria”. As a hybrid it cannot be placed in any series, and chromosome counts are not practical, so: n=? Kimnach decided it is important enough to be listed formally in the 2003 Lexicon, but with full explanation. He described it as 6 cm tall with leaves green not glaucous, the infl. cymose, 3 branched to 20 cm tall, flowers scarlet. It is a nice plant in cultivation.

=*Ech. x pulchella (Berger, 1904), current, but Berger only proposed it as a species & did not formally publish it, so the name is not official.

PULIDONIS (Walther, 1972), E. Cent. Mexico in state of Hidalgo, honors Sr. Miguel Pulido (an early Mexican Echeveria collector who probably was first to collect it in the wild); this is an attractive small species with milky green leaves having a red edge; plus pretty, pure yellow flowers. Walther put it in his old series 2 (Secundae) sp. #23, but later moved to Uhl’s revised Series Urceolatae in Haseltonia # 4 on p. 69; then in the 2003 Lexicon Kimnach places it in the new Series 16 (Urbiniae); photo in Walther’s book on p. 122 (fig. 60, p. 123), pub. in Ech. 1972, p. 122, see also Amer. Journ. Bot. 1992, p.561; Chromosome counts of n=32, 62 & 64 have been recorded (the 64 is an auto-tetraploid). Plants are nearly stemless, no hairs, little branching; leaves green with bright red margins; the infl. a cincinnus, nodding (down-curved tips) 18 cm tall, fl. yellow. Popular: found at most nurseries.

=*Ech. pulidonis [Walther, 1972], current.

 

PULVINATA (Rose, 1903), SW Mexico in state of Oaxaca; name means “pillow shaped” from Latin “pulvin” for cushion; published in Bulletin NY Bot. Gard, 1903. III: 5. This is one of the “fuzzy leaved Echeverias” considered highly attractive, especially in the forms with reddish-hirsute leaves, ‘Ruby’. Alfred Lau found a "near pulvinata", Lau-014, mentioned by Kimnach in the 2003 Lexicon as the attractive whitehaired form from Oaxaca, ‘Frosty’. Kimnach now places this species in the new Series 4 (Echeveria) in the 2003 Lexicon, with the cytology shown as being n=23. Normal plants are entirely hirsute except inside the flowers! The loose rosettes are on a branching 5 inch stem; the thick leaves are green, but densely covered with white or reddish hair. The inflorescences are between spikes and racemes in form, even subpaniculate below, 12 cm tall, flowers red.

=Cotyledon pulvinata [Hooker, 18??]. obs.

=*Ech. pulvinata [Rose, 1903], current.

 

 

 

pumila (Van Houtte, 1846), Cent. Mexico, Walther series 2, sp. #30, p. 138; on p. 35 of Haseltonia #3 of 1995 Uhl reduced this sp. to synonomy with Ech secunda var. glauca (30b). For details see under Ech. secunda in this listing, and in 2003 Lexicon where Kimnach reduced this to a mere un-named form of Ech. secunda type-species. [Note: some people will not understand why it has been so often reduced and combined. The reason has to do with the many forms of this species forming a “spectrum” in which it becomes hard to say where to place any one. In all such cases today botanists just combine the many forms into a highly variable single species. The extreme forms in such a spectrum can look very different from each other, and it is worth while to grow several of these forms. It is proper to call it by the new combined species name, with the form it had previously been known as added as a “forma” to identify it correctly].

=Cotyledon glauca (Baker, 1869), obs. This form found near Mexico City on volcano Popocatepetl & a bit into Puebla state.

=Ech. pumila var. glauca (Walther, 1972), obs. For this form see photo in CSSA Journ. Nov. 1992, p.303, Walther #30b.

=Ech. secunda var. glauca (Uhl, 1995), obs.

=*Ech. secunda [Booth, 1838], current.  Note: for outstanding named forms, just add that name after this one as a “forma”.

pumilglauca, (name of uncertain origin, probably a commercial cultivar name for one, or maybe all pumila forms).

PURPUSORUM (Berger, 1930), E. Cent. to SW Mexico in states of Puebla to Oaxaca in Sierra Mixteca mtn. range; name honors Joseph Purpus, who with brother Carl collected plants in Mexico (this species was collected with both working together); this species is attractive and distinct due to the fact that its leaves have an unusual dark mottling; first placed into old Walther series 2, sp. #24, p. 125; later moved into Uhl’s revised Series Urceolatae on p. 69 of Haseltonia #4, but in the 2003 Lexicon Kimnach now lists this in the new Series 16 (Urbiniae); ISI offering in CSSA Journ. Mar. 1978, p. 91 as ISI-1077, see Haseltonia #4 of 1996 on p. 85;  cytology: n=27 (diploid). The plants rarely offset, so propagation is from leaves, but is easy. The leaves are a light olive-green, heavily mottled & spotted by irregular markings of reddish-brown color, margins translucent; the inflorescences are unbranched cincinni to 20 cm tall; flowers rose below, scarlet above and yellow at the tips.

=Urbinia purpusii (Rose, 1911), obs.

=*Ech. purpusorum (Berger, 1930), current.