ECHEVERIA LISTINGS S

By James E. Low

sanchez-mejoradae (Walther, 1972), E. Cent. Mexico in state of Hidalgo; grows on canyon walls, first placed in old Walther series 2, sp.#17, p. 108, (fig. 48, p.109), honors a Mexican C&S botanist; published in book ‘Echev.’ 1972, p. 108 sp. First in series Secundae, but in Haseltonia #4 of 1996 Uhl wrote: “very similar to Ech. halbingeri.” Uhl moved it into Series Urceolatae as new field collected plants had n=60-62 (tetraploid), with one 10-ploid at n=150 (plus or minus 10), Kimnach took action on this in 2003 Lexicon and moved it again, this time into the new Series 16 (Urbiniae).

=Ech. sanchez-mejoradae [Walther, 1972], obs.

=*Ech. halbingeri var. sanchez-mejoradae [Kimnach, 1997], current.  This was a Comb. Nova in Haseltonia #5, p. 51. See halbingeri listing for details.

x SAYULENSIS (Walther, 1972), W. Cent. Mexico in state of Jalisco: Walther said it was from near Sayula (thus the species name); published in book ‘Echev.’ 1972. p. 151. Prof. Uhl in CSSA Journ 1989:6 wrote that it is a natural hybrid. Suspecting it to be a hybrid Uhl did an interesting procedure to prove it (see details in 2003 Lexicon). See BW photos & drawing in CSSA Journ. Nov. 1975, figs.4, 5, & 6 on p. 265. Kimnach does not try to place hybrids in any series, & usually chromosome counts are not practical for hybrids, but for this one he lists the cytology of n=43, and his description includes these points: short, branching stems, rosettes to 25 cm in diameter, leaves green and slightly glaucous, inflorescences are panicles to 36 cm tall, and flowers pink.

=*Ech. x sayulensis [Walther, 1972], current.

SCHAFFNERI (Rose, 1903), NE to E Cent. Mexico in states of NL, SLP to Puebla, grows on gravel/sand outwashes at foot of mountains and sometimes on flat, dry land; republished in Bulletin NY Bot. Gard, 1903, III: 9; now in new Series 1(Angulatae) in 2003 Lexicon; cytology: n=12. Refer to Haseltonia #6, p. 65 & 81 for discussion. Plants rarely branching, rosettes of green to purplish leaves not glaucous; the inflorescences of 2 cincinni to 30 cm tall, flowers yellow and pink.

=Cotyledon schaffneri [Watson, 1882], obs.

=Ech. teretifolia var. schaffneri [Walther, 1935], obs.

=*Ech. schaffneri [Rose, 1903], current.

SCHEERI (Lindley, 1845), SW Mexico in state of Oaxaca, probably near village of Chontal; first listed in old Walther series 2, #40, p. 154, but now listed in new Series 5 (Gibbiflorae); published in Edw. Botanical Reg. 1845, 31:27; species named to honor Frederick Scheer of Kew, England, a succulent plant collector in mid-1800s. Plants rarely branching, lower stems to 10 cm; medium rosettes of glaucous green leaves with red margins; inflorescences cymose, 2 or 3 branched, short; flowers orange to pink. Walther wrote that this species was discovered by MacDougall, but it is not in his notes. Oddly Kimnach does not give the chromosome count in the 2003 Lexicon, probably an oversight.

=Cotyledon scheeri, [Baker, 1869], obs.

=*Ech. scheeri, [Lindley, 1845], current.

SECUNDA (Booth, 1838), E. Cent. Mexico in state of Hidalgo, etc. in mts., a variable and widespread species, name refers to “up pointing” (for flowers), see CSSA Journ. Jul. 1994, p. 172, first placed in old series Secundae, and in 2003 Lexicon it is in new Series 13 of same name; cytology: n=30, 32 and it is a tetraploid, see in AJB of 1982, p. 846. Also see Haseltonia #3 of 1995 on p. 34 & 35. Uhl combines some old species into Ech. secunda (listed below). Above species is central sp. for Series Secundae, mostly tetraploid. (the revised Uhl Series Secundae; to have only 4 valid species). Note: The common form of this species is probably the best starting plant for a beginning collector. Also note that this is one of the most commonly planted of all Ech. species over the world as a bedding plant, or in mild climates as a rock garden plant. The plants are medium in size, freely branching and form nice rosettes of blue-glaucous leaves, inflorescences are a cincinnus 5 to 15 cm tall, and flower colors range from orange-red through deep pinks. This species is widespread in east central Mexico and it adapts to many habitat types, even high altitude. It is also extremely variable over its range, thus accounting for the fact that in earlier times many were thought to be full species and were published as such; but later study found them to form a “spectrum”, and the only practical way to handle that situation in modern botany is to combine all under the name of one highly variable central species, sometimes giving the most extreme forms the status of “forms” or “variety”. The reason for such combinations is that it is impossible for any expert to decide the identifications of the many intermediate forms. This certainly does not say that the variations are not real, or not much different from each other: the differences are real, and good collections should indeed include a range of types. These plants are of easy culture and attractive, thus popular. In the 2003 Lexicon 2 extreme forms are formally recognized as valid “forms” for this species today; but some of the others are in reality forms as well and can be so designated on their name tags (for example: “Ech. secunda fma. cornuta”). The list of synonyms is rather long:

=Ech. secunda, (Booth, 1838), current: still valid name today as the central species to the spectrum of many local types, some known as named forms, others not even named.

=Cotyledon secunda [Baker, 1869], obs.

=Ech. alpina [Walther, 1935], obs. Valid as a form.

=Ech. cornuta [Walther, 1972], obs.        “

=Ech. elatior [Walther, 1935], obs.           “

=Ech. pumila [Van Houtte, 1846], obs.     “

=Cotyledon pumila [Baker, 1869], obs.

=Ech. s. var. pumila [Otto, 1873], obs.

=Ech. s. var. pumila [Poellnitz, 1936], obs.

=Ech. s. var. glauca [Otto, 1873], obs.

=Ech. spilota [Kuntze, 1853], obs.

=Cotyledon glauca [Otto, 1873], obs.

=Ech. s. var. glauca [Otto, 1873], obs.

=Ech. reglensis [Walther, 1972], obs.

=Ech. glauca [Morren, 1874], obs.

=Ech. pumila var. glauca [Walther, 1972], obs.

+*Ech. sec. forma byrnesii [Kimnach, 1997], current: (Kimnach formalizes Uhl’s above sugg. in Haseltonia #5, p. 51). This is an alpine (mountain) form, distinctive because of its plain green leaves (the others have blue-glaucous leaves).

+*Ech. secunda forma secunda [Booth, 1835], current. (the type-species).

sedoides (Walther, 1958), SW Mexico in state of Oaxaca, name refers to (a plant) "resembling a Sedum", pub. in CSSA Journ. 1958, 30:153, (the flowers do resemble those of a Sedum, but plant is an Ech.), Series 8 (Nudae)., n=19 by Uhl in 1982. Sedoides is now combined into another species:

=Ech. sedoides, [Walther,1958], obs.

=*Ech. macdougallii [Walther, 1958], current.

SEMIVESTITA (Moran, 1954), of NE Mexico in 5 states including SLP, NL & Hidalgo; first placed in the old Walther series 2, sp. #45, p. 166, now (in 2003 Lexicon) tentatively placed in new Series 5 (Gibbiflorae) as there is some doubt about this series for this plant. The blueish sepals are unique to this species; name  means “semi-clothed”, published in CSSA Journal 1954, 26:60; cytology: n=17; lower stems 10 to 15 cm and about 1 cm in dia. rarely branching, leaves hairless to minutely pubescent, green, margins purple-red; inflorescences paniculate and up to 55 cm tall with several branches, sepals green to purplish or blueish-glaucous & pruinose, flowers pink to red; 2 varities currently recognized:

+*var. semivestita [Moran, 1954], current. CSSA Journ. '54, 26:6, #45a. Type-species.

+*var. floresiana [Walther, 1958], current. Santa Ana at 1585 M. elev., pub in CSSA Journ. '58, 30:109, Walther sp. #45b.

=*Ech. semivestita [Moran, 1954], current. Type-species.

SESSILIFLORA (Rose, 1905), S. Mexico in states of Chiapas & Oaxaca near Mitla at 2200 m alt., name means “flowers without stem”; first placed in old Walther series 2, sp. #45, p. 166, but in the 2003 Lexicon Kimnach places this species in the new Series 7 (Mucronatae); cytology: n=15 in AJB of 1982, p. 846, and in new Lexicon. Plants branch & have medium rosettes, leaves very glaucous & pale green with brown-purple margins, the inflorescence a medium spike 15 to 30, rarely as much as 40 cm tall, flowers coral-red.

=Ech. corallina [Alexander, 1941], obs.

=*Ech sessiliflora [Rose, 1905], current.

SETOSA (Rose & Purpus, 1910), E. Cent. Mexico in states of Puebla & Oaxaca. This species is so extremely varied that today 5 varities are recognized in the 2003 Lexicon, and intermediates also exist; name means “bristled”, so this species is another of the “fuzzy Echeverias”; see CSSA Journ. of Jan. 1993, p. 27 by Moran for discussion of all var. listed below, all diploids; this species was published in Contributions US Nat. Herbarium, 1910; first placed in old series Ciliatae, and in the 2003 Lexicon it is in the new Series 3 of the same name; cytology: n=25 (diploid), see Haseltonia #5, p. 21 & 22. Plants of the type-species have main stems of about 10 cm long by 1 to 2 cm dia. and branch freely forming several rosettes; leaves are green and are hirstute & ciliate with hairs of 2 to 3 mm, the inflorescences are a hirsute cincinnus to 30 cm tall with about 10 fl.; flowers are orange-red below to yellow near tip.

=*Ech. setosa [Rose & Purpus, 1910], current.

=*var. setosa [Rose & Purpus, 1910], current: type species.

+*var. ciliata [Moran, 1993], current: CSSA Journ. '93, 65:I: 32, Oaxaca, photo p. 99 CSSA of Mar. 1995, differs from type-sp. in minor ways, often more flowers in inflorescence.

=Ech. ciliata [Moran, 1961], obs.

+*var. deminuta [Meyran, 1989], current, pub. in Mex. C&S Journ. 1989, XXXIV, 4:75; a heavily branching type (mat-forming), few hairs, leaves green, or even glaucous.

=Ech. ‚Jim Rundell’ =Ech. rondelii, [Sterk, 1989], obs. (names invalid). Hort type.

+*var. oteroi [Moran, 1993], current:  CSSA Journ. 1993, 65, I: 33, Oaxaca. A sparsely branching type with mostly hair free leaves (not fuzzy!).

+*var. minor [Moran, 1993], current: CSSA Journ, 1993, 65(1). p 33-35; a dwarf form found by Father Fittkau.

(note: varities ciliata & minor were offered by ISI in CSSA Journ. Mar.  95, p.99: ciliata was ISI 95-28, & minor was ISI 95-29).

 

 

 

E. ‘Set-Oliver’ is a cultivar listed by Walther (CSJA 8: 172, 1937) and still popular in nurseries today. It is a hybrid by V. Reiter with parents being E. setosa + E. harmsii.

SHAVIANA (Walther, 1972), NE to Cent. Mexico in states of NL, Tamaulipas, & Coahuila where it grows in shade in pine needles, see CSSA Journal Jul. 1994 on p. 172 & p. 178, color photo cover of CSSA Journ. Nov. 1993, name was for the Missouri Botanical Garden nicknamed "Shaw's Garden" (its botanists were the first to collect this sp.) pub. In book Ech. 1972, p. 270; BW in-habitat photo CSSA Journ. Nov. 1975, p. 266; first in old series 7 (Pruinosae), now in new Series 1 (Angulatae), cytology: n=13, see Haseltonia #5, p. 33, and refer to Haseltonia #6, p. 81. Plants are hairless, little branching, rosettes 4 to 5 inch, thin leaves of glaucous greenish to pinkish, whitish margins undulate-crispate, inflorescences 1 to 2 cincinni to 30 cm tall, flowers rose colored. Only slowly offsetting.

= *Ech. shaviana [Walther, 1972], current.

SIMULANS (Rose, 1905), NE Mexico in state of NL, see table 1, p. 68 of Haseltonia #4 of 1996, and text on p.85; this was in Uhl’s revised series Urceolatae, but in 2003 Lexicon it has been placed in the new Series 16 (Urbiniae), cytology: n=32 (diploid). The Latin simulans means “similar to” in reference to how much this species resembles Ech.elegans. Published in N. Amer. Flora 22(1): 22, 1905. This is a slowly branching type with large, loose rosettes of glaucous, greenish-white leaves, the inflorescence has 1 or 2 cincinni from 20 to 40 cm tall, flowers rose-pink to yellowish at tips. This species differs from the many forms of Ech. elegans by having larger and looser rosettes, leaves and corollas.

=Ech. elegans ssp. simulans (Poellnitz, 1936), obs.

=*Ech. simulans (Rose, 1905), current.

 

SKINNERI (Walther, 1972), SW Mexico in state of Oaxaca in Cerro Madrena mts. at 7000 ft. elev., pub. in Ech.1972, p.307. Type has MacDougall field collection number of B-204. In the 2003 Lexicon it is listed in the new Series 8 (Nudae) with cytology of n= (about) 40. Plants have long lower stems 30 or more cm. tall, freely branching; leaves scattered and in loose rosettes, smooth green often with reddish margins, the inflorescence a raceme from 15 to 20 cm tall, flowers scarlet with yellow margins.

=*Ech. skinneri [Walther, 1972], current.

 

spathulifolia (de Smet, 18??), This “Echeveria” was in reality a Sedum!

=Ech. spathulifolia [de Smet, 18??], obs.

=Sedum spathulifoliu, (Hooker, 1834) current.

SPECTABILIS (Alexander, 1941) SW Mexico in state of Oaxaca in Sierra Juaraz, means “remarkable” (referring to the plant with its large, attractive flowers), pub. in CSSA Journ. 1941, XIII: 137; was in old Series 9 (Nudae), now in 2003 Lexicon in new Series 8 of same name, cytology: n=ca. 105. Read page 303 of CSSA Journ. of Nov. 1999. Lower stems can be quite tall (up to 60 cm) and branching, loose rosettes of green leaves with red margins; the inflorescences are tall racemes to 70 cm with large flowers of scarlet to orange and yellow margined.

=*Ech. spectabilis [Alexander, 1941], current.

sprucei (Berger, 1930), Ecuador near Ambata; pub. in Engl. Nat. Pfl. fam. 1930, II: 18a: 473; new Series Nudae, n=22. This sp. now combined into a different central species.

=Cotyledon sprucei (Baker, 1869), obs.    Pub. In Belgique Hortic, XXIV, p. 167.

=Ech. sprucei (Berger, 1930), obs.

=*Ech. quitensis var. sprucei [Poellnitz, 1936], current.

STEYERMARKII (Standley, 1944), N. Guatemala at 1300 to 3700 m elev.; first placed in Walther series 2, #43, p. 163, pub. in Field Museum Bot. Ser., 1944, 23:4:16. Now in new Series 5 (Gibbiflorae) in the 2003 Lexicon; Note: this is a little-known species with no illustrations, and is not now in cultivation anywhere, thus the cytology is unknown: n=? It was said to grow epiphytic on rocks at upper altitudes, the following description is on record: stems 5 cm tall by 1 cm dia. usually unbranched, leaves rosulate and green tinged purplish or rose; the inflorescences 2 branched racemes, 5 to 20 cm long with few flowers, the flowers red to yellowish with yellow margins.

=*Ech. steyermarkii [Standley, 1944], current.

STOLONIFERA (Otto, 1873), Mexican habitat unknown (or this may be a hort type from culitv.), Walther placed it in series 2, #39, p. 152, but in 2003 Lexicon Kimnach merely shows a question mark for the series: it is possibly a hybrid of Ech. glauca and Ech. grandifolia. Pub. in Hamburger Gartenzeitung, 1873, XXIX:9, (the name means “bearing stolons or runners”). No cytological count is given anywhere, thus: n=? Note: this possible species was known since early times (1869), but since it cannot be found in nature, questions arose as to its true status. Walther found garden hybrids that seemed to be this same plant, and he thought the parents were Ech. glauca X grandifolia. Nevertheless, it is popular even today, and it is easy and quite attractive. It deserves recognition. It is a neat branching plant with greyish green rosette leaves, and medium secund inflorescences and reddish flowers.

=Cotyledon stolonifera (Baker, 1869), obs.

=*Ech. stolonifera (Otto, 1873), current.

STRICTIFLORA (Gray, 1852), N. Cent. Mexico & a bit into S. Texas, species name means “flowers tight” (almost closed), pub. in Plant Wright., 1850, I:76, see Haseltonia #3 of 1995 on p. 28 where Uhl only mentions this sp.; cytology: n=12: refer to Haseltonia #6, p. 66 & 83. Short stems, plants rarely branching, thin rosette leaves of grey-green to buff-green, sometimes with red margins; the inflorescences are second racemes to 25 cm tall, flower base rose to yellow or reddish above. Plants said to be difficult to maintain in cultivation. This is the only species in this genus whose range comes even a tiny bit into the USA, maybe 3 miles at one spot. Some forms from Mexico have wider leaves and may become new species after a period of study.

=Cotyledon strictiflora [Baker, 1869], obs.

=*Ech. strictiflora [Gray, 1852], current.

 

 

SUBALPINA (Rose & Purpus, 1910), E. Cent. Mexico in Vera Cruz state to Puebla state, on lava 2500 m elev.; name “subalpina” means the species grows at intermediate mtn. altitudes; first placed in old Walther series 2, #36, p.148, now listed in new Series 13 (Secundae) in 2003 Lexicon; pub. in Contributions to US Nat. Herbarium 1910, 13:45; see Haseltonia #3 of 1995, p. 37 for Uhl’s update of this sp.; cytology: n=34 (tetraploid). This is a rarely branching species with rosettes of glaucous green leaves, the inflorescence a 2-branched cincinnus to 30 cm. tall, flowers orange to red with yellow margins. The wild plants a bit variable in rosette details.

=Ech. akontiophylla [Werdermann, 1932], obs.

=Ech. meyraniana [Walther, 1959], obs. (combined into subalpina by Uhl in 1995).

=*Ech. subalpina, [Rose & Purpus, 1910], current.

Pour voir plus de photos/to see more pictures

 

 

SUBCORYMBOSA (Kimnach & Moran, 1994), SW Mexico in state of Oaxaca; name refers to flower clusters (short coymbose raceme, long pedicels), pub. in CSSA Journ. Jan. 1994, p. 11; ISI in CSSA Journ. Mar. 1993, p. 74, (ISI 93-32); now in new Series Racemosae, in 2003 Lexicon; cytology: n=29 (a most unusual count). Small rosettes of glaucous, blueish-grey leaves, with reddish margins and often flecked red below; the inflorescences are short racemes appearing corymbose, flowers reddish-orange below to yellow upper. Note: a wider leaf form has been found that after study could become a variety (Lau 026). Color photo on p. 198 & 199 of CSSA Journ. of Sept. 2007.

=*Ech. subcorymbosa [Kimnach & Moran, 1994 (Lau 030), current.

 

 

SUBRIGIDA (Rose, 1903), E. Cent. Mexico in 4 states growing on cliffs, name means “somewhat rigid”, first placed in old Walther series 2 (Secundae), sp. #46; [see in CSSA Journ., Jul. 1979, p.178 about an imposter], & in CSSA Journ. Mar. 1976, p. 65 (BW photo & a few words); pub. in Bulletin NY Bot. Gard, 1903, III: 10. Photo & text CSSA Journ. p. 241-3, Sept. 1997; now in new Series 5 (Gibbiflorae) in 2003 Lexicon; cytology n=27or 54; plants rarely branching, medium rosettes; leaves white-pruinose or green, red margins, edges often finely undulate-crenulate, the inflorescences are tall cymes to one meter, flowers glaucous pink to scarlet; an attractive species.

=Cotyledon subrigida [Robinson, & Seaton, 1893], obs.

=Ech. palmeri [Rose, 1903], obs.

=Ech. rosei [Nelson & Macbride, 1913], obs.

=Ech. angusta [Poellnitz, 1936], obs.

=*Ech. subrigida [Rose, 1903], current.

 

subsessilis (Rose, 1905), E. Cent. to SW Mexico in states of Puebla to Oaxaca in Sierra Mexteca Mts., name refers to individual flowers “almost stemless”, ISI-1200: CSSA Journ. Mar., 1980, p. 101, pub. N. Amer. Flora, 1905, XXII:20; see in Haseltonia #6, p. 64. This sp. now combined; see below.

=Ech. subsessilis [Rose, 1905], obs.

=Ech. desmetiana [Morren, 1874], obs.

=*Ech. peacockii [Croucher, 1874], current.

subspicata (Berger, 1930), is now combined into Ech. bicolor, see CSSA Journ. of Jan. 1992, p. 3 & 4, for all details.

=Cotyledon subspicata (?) [Baker, 1869], obs.

=Ech. subspicata [Berger, 1930], obs.

=*Ech. bicolor [Walther, 1935], current.